RDA
RDA_vignette.RmdRedundancy analysis (RDA)
#Install required packages
rda_packages()If using RDA, please cite the following: Capblancq T., Forester B.R. (2021) Redundancy analysis: A Swiss Army Knife for landscape genomics. Methods in Ecology and Evolution 12:2298-2309. DOI:10.1111/2041-210X.13722
Redundancy analysis (RDA) is a genotype-environment association (GEA) method that uses constrained ordination to detect outlier loci that are significantly associated with environmental variables. It does so by combining regression (in which response variables are genetic data, in our case, while explanatory variables are environmental data) with ordination (a PCA). Importantly, RDA is multivariate (i.e., multiple loci can be considered at a time), which is appealing in cases where multilocus selection may be occurring.
Model selection is performed by starting with a null model wherein the response is only explained by an intercept, and environmental variables are added until the amount of variance explained by a model that includes all variables is reached. By doing so, this method minimizes the redundancy among explanatory variables. Importantly, we can also perform an RDA that accounts for covariables (or “conditioning variables”) such as those that account for neutral population structure. When covariables are included, this is called a partial RDA. In algatr, population structure is quantified by running a PCA on the genetic data, and selecting a number of PC axes to best represent this type of neutral variation.
Some of the earliest examples of uses of RDA for identifying environment-associated loci are Lasky et al. 2012 and Forester et al. 2016. Since then, there have been several reviews and walkthroughs of the method that provide additional information, including workflows and comparisons with other GEA methods (e.g., Forester et al. 2018). Much of algatr’s code is adapted from one such paper, Capblancq & Forester 2021, the code of which is available here. algatr’s general workflow roughly follows the framework of Capblancq & Forester (2021).
RDA cannot take in missing values. Imputation based on the per-site
median is commonly performed, but there are several other ways
researchers can deal with missing values. For example, algatr contains
the str_impute() function to impute missing values based on
population structure using the LEA::impute() function.
However, here, we’ll opt to impute to the median, but strongly urge
researchers to use extreme caution when using this form of simplistic
imputation. We mainly provide code to impute on the median for testing
datasets and highly discourage its use in further analyses (please use
str_impute() instead!).
The general workflow to perform an RDA with algatr is as follows:
Simple vs. partial RDAs: We can run a simple RDA (no covariables) or partial RDA (conditioning covariables included) using
rda_run()Variable selection: Both simple and partial RDAs can be performed considering all variables (
"full"model) or by performing variable selection to determine variables that contribute most to genetic variance ("best"model)Variance partitioning: We can perform variance partitioning considering only variables that contribute significantly to genetic variance using
rda_varpart()Detecting outlier loci: Based on the results from above, we can detect outlier loci with
rda_getoutliers()
Read in and process genetic data
Running an RDA requires two data files for input: a genotype dosage
matrix (the gen argument) and the environmental values
extracted at sampling coordinates (the env argument). Let’s
first convert our vcf to a dosage matrix using the
vcf_to_dosage() function. N.B.: our code assumes that
sample IDs from our genetic data and our coords are in the same order;
be sure to check this before moving forward!
load_algatr_example()
#>
#> ---------------- example dataset ----------------
#>
#> Objects loaded:
#> *liz_vcf* vcfR object (1000 loci x 53 samples)
#> *liz_gendist* genetic distance matrix (Plink Distance)
#> *liz_coords* dataframe with x and y coordinates
#> *CA_env* RasterStack with example environmental layers
#>
#> -------------------------------------------------
#>
#>
# Convert from vcf to dosage matrix:
gen <- vcf_to_dosage(liz_vcf)
#> Loading required namespace: vcfR
#> Loading required namespace: adegenetAs mentioned above, running an RDA requires that your genotype matrix contains no missing values. Let’s impute missing values based on the per-site median. N.B.: this type of simplistic imputation is strongly not recommended for downstream analyses and is used here for example’s sake!
# Are there NAs in the data?
gen[1:5, 1:5]
#> Locus_10 Locus_15 Locus_22 Locus_28 Locus_32
#> ALT3 0 0 0 0 0
#> BAR360 0 0 0 NA 0
#> BLL5 0 NA 0 0 NA
#> BNT5 0 0 NA NA 0
#> BOF1 0 NA NA NA NA
gen <- simple_impute(gen)
# Check that NAs are gone
gen[1:5, 1:5]
#> Locus_10 Locus_15 Locus_22 Locus_28 Locus_32
#> ALT3 0 0 0 0 0
#> BAR360 0 0 0 0 0
#> BLL5 0 0 0 0 0
#> BNT5 0 0 0 0 0
#> BOF1 0 0 0 0 0Process environmental data
Let’s extract environmental variables using the
extract() function from the raster package. We also need to
standardize environmental variables. This is particularly important if
we are using (for example) bioclimatic variables as input, as units of
measurement are completely different (e.g., mm for precipitation
vs. degrees Celsius for temperature). To do so, we’ll use the
scale() function within the raster package.
# Extract environmental vars
env <- raster::extract(CA_env, liz_coords)
# Standardize environmental variables and make into dataframe
env <- scale(env, center = TRUE, scale = TRUE)
env <- data.frame(env)Running simple and partial RDAs using rda_run(), with
and without variable selection
The main function within algatr to perform an RDA is
rda_run(), which uses the rda() function
within the vegan package.
Run a simple RDA with no variable selection
A simple RDA is one in which our model will not account for
covariables in the model, and is the default for rda_run().
Let’s first run a simple RDA on all environmental variables (i.e., no
variable selection). This is specified using the
model = "full" argument.
mod_full <- rda_run(gen, env, model = "full")The resulting object is large, containing 12 elements relevant to the RDA. Let’s take a look at what function was called. We can see that all environmental variables (CA_rPCA1, 2, and 3) were included in the model, and that geography or structure were not included in the model (and there were no conditioning variables).
mod_full$call
#> rda(formula = gen ~ CA_rPCA1 + CA_rPCA2 + CA_rPCA3, data = moddf)Now, let’s take a look at the summary of this model. One of the most important parts of this object is the partitioning of variance. Within an RDA, the term “inertia” can be interpreted as variance, and our results show us that the amount of variance explained by explanatory variables alone (constrained inertia) is only 9.768%. Unconstrained inertia is the residual variance, which is very high (90.232%). The summary also provides site scores, site constraints, and biplot scores.
head(summary(mod_full))
#> $species
#> RDA1 RDA2 RDA3 PC1
#> Locus_10 3.072512e-02 -1.130156e-02 2.568722e-02 -1.786256e-02
#> Locus_15 8.232018e-02 -5.560180e-02 -9.948805e-02 -9.372926e-02
#> Locus_22 9.597919e-03 1.021082e-03 2.333238e-02 1.249210e-01
#> Locus_28 -7.716309e-02 -8.271105e-02 -6.967672e-02 -2.229286e-02
#> Locus_32 1.095373e-02 -1.691960e-04 3.178787e-02 -1.579991e-02
#> Locus_35 -4.633795e-32 -1.047216e-31 -8.297790e-32 -6.887286e-17
#> Locus_37 -6.502873e-33 3.993792e-32 -1.706487e-32 -5.800186e-17
#> Locus_61 5.337664e-02 -9.209774e-02 4.866692e-03 7.297376e-02
#> Locus_71 1.386618e-03 5.808684e-03 1.769149e-02 1.004786e-01
#> Locus_72 1.222591e-32 1.540195e-32 7.673218e-32 2.777891e-17
#> Locus_82 1.121917e-32 6.524345e-33 3.587319e-32 -9.477760e-17
#> Locus_84 -1.282353e-02 2.668403e-02 -1.607300e-03 4.364401e-02
#> Locus_90 5.097845e-02 -2.527629e-02 1.165401e-01 -7.989586e-02
#> Locus_91 8.484251e-02 -7.203143e-02 9.442310e-03 -3.381549e-02
#> Locus_96 2.233926e-35 5.099313e-33 -1.524506e-32 5.005347e-17
#> Locus_104 -1.986935e-02 -1.482855e-02 5.189881e-03 -9.515613e-03
#> Locus_122 4.248706e-02 -8.344988e-03 -2.912039e-02 9.955968e-02
#> Locus_125 1.073239e-01 -7.623381e-02 3.729670e-02 -2.123165e-02
#> Locus_131 -7.877379e-03 1.564754e-02 2.114341e-02 5.007196e-02
#> Locus_133 1.378087e-33 1.680148e-33 4.945054e-33 2.129458e-17
#> Locus_138 1.091500e-01 1.997988e-02 1.347650e-02 -6.378991e-02
#> Locus_154 -3.296383e-02 -1.597652e-02 3.560521e-02 6.852150e-02
#> Locus_162 3.267219e-02 -1.175728e-01 5.398876e-02 -7.614820e-02
#> Locus_166 1.413161e-01 -1.030291e-01 4.777541e-02 -7.024082e-02
#> Locus_177 5.401464e-34 -4.750549e-33 1.220190e-33 -5.920464e-17
#> Locus_185 4.856833e-34 7.233246e-33 -3.042676e-33 -3.844501e-17
#> Locus_191 1.169980e-02 1.036416e-01 -4.703122e-02 -3.337053e-02
#> Locus_193 -4.908999e-02 -2.124439e-02 -1.820482e-02 -1.346309e-01
#> Locus_207 3.261014e-02 1.005953e-02 -4.106132e-02 -1.779306e-02
#> Locus_211 2.537180e-02 -2.364704e-02 2.817155e-03 -1.373421e-02
#> Locus_213 -5.188078e-02 -1.627408e-02 -6.265149e-02 -4.945625e-03
#> Locus_214 4.962746e-02 -1.021595e-01 -7.312763e-02 -2.991293e-02
#> Locus_222 -1.408912e-02 3.117436e-02 8.811572e-03 3.524181e-02
#> Locus_227 1.346156e-01 -1.098083e-02 1.257735e-02 -1.670522e-01
#> Locus_242 4.587131e-03 9.233846e-03 3.083404e-03 -7.822608e-03
#> Locus_249 1.680963e-01 4.845511e-02 -4.373299e-02 -9.653170e-02
#> Locus_254 -4.414339e-02 -1.283360e-02 6.089062e-03 -1.883158e-02
#> Locus_256 2.482736e-02 2.072786e-02 8.431861e-02 2.297250e-02
#> Locus_263 1.029768e-01 -1.967721e-01 -2.974146e-03 -1.144217e-01
#> Locus_268 1.547392e-03 -1.550198e-02 -1.668626e-02 5.767993e-02
#> Locus_279 -7.788026e-03 -9.177332e-04 -5.354875e-02 -1.694952e-02
#> Locus_285 1.848936e-02 -6.298659e-02 5.505993e-02 -7.140754e-02
#> Locus_287 -6.636351e-02 -3.393219e-02 -1.483964e-02 -1.230938e-02
#> Locus_292 -7.642827e-04 2.549682e-03 -1.685845e-02 5.847707e-02
#> Locus_294 -6.183308e-03 5.346666e-02 1.585868e-02 1.640659e-02
#> Locus_303 2.083966e-34 1.061950e-33 1.240101e-34 -2.622149e-17
#> Locus_308 -1.204379e-02 2.102857e-02 3.709140e-02 -1.975608e-02
#> Locus_312 -9.585422e-03 1.313767e-01 -5.412095e-02 1.227400e-03
#> Locus_313 1.106332e-34 2.214406e-34 2.376467e-35 -9.867672e-17
#> Locus_314 5.813472e-02 3.980658e-02 1.000765e-02 -2.943692e-02
#> Locus_317 -1.291548e-02 1.901979e-02 1.703403e-02 6.323903e-02
#> Locus_318 -3.993283e-02 -1.730491e-02 4.357889e-02 -1.765347e-02
#> Locus_323 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_330 -8.333460e-04 -1.670290e-02 -3.339391e-03 6.629651e-02
#> Locus_331 2.134327e-02 7.225468e-03 -3.332978e-02 -4.327155e-03
#> Locus_333 1.551458e-02 -2.395792e-02 1.155128e-02 -1.658104e-02
#> Locus_334 1.974546e-02 -3.838672e-03 7.812471e-03 -6.920496e-03
#> Locus_336 -1.494260e-02 -4.583701e-03 3.575609e-03 -3.258059e-03
#> Locus_337 4.203697e-02 -2.892541e-02 -3.376534e-03 -1.748853e-02
#> Locus_339 1.436718e-01 -2.346413e-02 -7.417133e-02 2.646974e-01
#> Locus_341 1.449424e-01 -1.093150e-01 -4.942921e-02 -6.792028e-02
#> Locus_345 4.338438e-03 8.116186e-02 -3.249917e-02 -8.813406e-04
#> Locus_350 1.332045e-01 -5.314949e-02 2.672613e-02 -2.300775e-02
#> Locus_354 1.933421e-02 7.817423e-02 -4.759152e-02 -1.936888e-02
#> Locus_359 9.988352e-02 1.613404e-02 -5.939488e-02 -2.886194e-01
#> Locus_365 1.225854e-02 8.366097e-03 -1.185795e-02 -1.051465e-02
#> Locus_366 1.446208e-02 -2.285700e-02 5.722978e-03 1.175648e-01
#> Locus_369 -5.763337e-02 -2.722022e-02 -1.953374e-02 -1.346475e-02
#> Locus_372 -4.344726e-04 4.361750e-02 -2.808839e-03 5.358313e-02
#> Locus_389 -4.627971e-02 -5.430740e-02 4.816851e-03 -6.578326e-02
#> Locus_391 3.538283e-02 -2.618367e-02 -3.175564e-03 -1.727891e-02
#> Locus_392 7.723314e-02 -8.337540e-02 -1.399527e-02 -4.781578e-02
#> Locus_393 -9.057143e-03 -1.138063e-02 1.370934e-02 6.248328e-02
#> Locus_394 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_396 3.379260e-02 -5.294546e-02 2.383823e-02 -2.474176e-02
#> Locus_399 7.917544e-02 -6.575207e-02 -8.072964e-03 -3.531519e-02
#> Locus_400 -8.222744e-03 4.948400e-02 -2.760177e-02 7.296207e-02
#> Locus_401 1.877062e-02 5.023278e-02 1.055960e-01 6.591692e-02
#> Locus_404 4.237154e-02 1.104944e-01 -8.342696e-02 -8.720722e-02
#> Locus_406 -2.844047e-01 1.167560e-01 1.463536e-02 -4.154901e-02
#> Locus_409 -4.253029e-02 -4.326751e-02 -2.253582e-02 -1.709879e-02
#> Locus_420 1.710223e-02 -2.460898e-02 3.111726e-02 9.462869e-02
#> Locus_422 -2.274937e-02 -1.711997e-02 3.632673e-02 1.227388e-01
#> Locus_427 5.212766e-02 6.551714e-02 -6.479956e-02 -6.719209e-02
#> Locus_430 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_436 -3.200534e-02 3.179776e-02 3.527449e-02 6.366290e-02
#> Locus_438 3.448262e-03 -2.667407e-02 -2.280999e-02 1.002383e-01
#> Locus_439 -1.160850e-02 4.610738e-02 3.518952e-02 -3.115790e-02
#> Locus_441 2.346099e-02 -4.436600e-02 3.117118e-04 -1.160622e-01
#> Locus_445 -1.301430e-02 2.041734e-03 1.731270e-02 2.396821e-02
#> Locus_448 1.561097e-02 2.256848e-02 1.812192e-02 1.026928e-01
#> Locus_449 1.973920e-02 3.848370e-02 -3.368024e-02 -3.287095e-02
#> Locus_456 -8.165593e-03 3.729787e-02 -1.538983e-02 1.198541e-01
#> Locus_457 -3.211412e-03 6.534026e-02 2.856715e-03 1.168707e-01
#> Locus_458 6.435961e-02 2.007850e-02 -3.289597e-03 3.424604e-02
#> Locus_460 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_461 -3.973869e-02 -2.965711e-02 1.037976e-02 -2.349882e-02
#> Locus_462 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_464 -5.183841e-02 -1.823147e-02 1.913700e-02 -3.860336e-02
#> Locus_469 1.231573e-01 -1.668373e-01 6.180401e-02 4.050704e-01
#> Locus_475 4.902874e-03 1.006275e-01 -4.227780e-02 -1.045867e-02
#> Locus_480 4.231170e-02 -5.608097e-02 -5.387688e-02 -8.108542e-02
#> Locus_482 -3.109519e-02 7.226611e-04 8.770170e-03 2.788869e-02
#> Locus_483 2.175061e-02 -2.415303e-02 -2.134472e-02 -1.936326e-02
#> Locus_491 -5.544226e-02 -2.833090e-02 -1.697848e-02 -1.046099e-02
#> Locus_498 1.085396e-03 1.262448e-04 3.739992e-02 2.321708e-01
#> Locus_500 1.900870e-03 -1.117209e-02 -6.123725e-03 3.572515e-02
#> Locus_503 -2.998202e-02 6.382269e-02 -1.322370e-02 5.288130e-02
#> Locus_505 -2.119874e-02 6.158320e-02 1.456022e-02 -5.063039e-02
#> Locus_508 1.297806e-02 8.278357e-02 -7.831450e-02 -1.298741e-01
#> Locus_512 8.049292e-02 -5.717536e-02 2.797253e-02 -1.473547e-02
#> Locus_516 3.819664e-03 -6.424957e-03 -1.189838e-02 -5.093542e-03
#> Locus_517 -2.233603e-02 2.134180e-01 -1.394747e-02 1.013401e-03
#> Locus_518 8.182019e-03 -1.774912e-03 1.813179e-02 2.325155e-01
#> Locus_528 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_529 5.568574e-02 -4.290147e-03 2.017993e-02 -2.535115e-02
#> Locus_531 2.800755e-02 -3.148967e-02 1.042443e-02 -3.867976e-02
#> Locus_532 5.594567e-03 1.046295e-02 3.997199e-02 -2.154311e-02
#> Locus_533 -5.093785e-02 -2.492084e-02 -1.471341e-02 -1.244150e-02
#> Locus_535 -1.384728e-02 3.423578e-02 2.709911e-02 -2.475132e-02
#> Locus_546 -2.229009e-02 3.171847e-02 3.269573e-02 3.539702e-02
#> Locus_557 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_558 8.165437e-02 3.781653e-02 3.256762e-02 3.110185e-02
#> Locus_561 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_562 -6.132617e-02 -4.775015e-02 7.277528e-03 -3.816071e-02
#> Locus_564 8.241669e-03 2.267897e-02 2.981431e-02 1.339624e-01
#> Locus_566 -1.474653e-02 7.463961e-02 -1.973038e-03 1.883996e-02
#> Locus_572 -1.865789e-02 -1.945389e-02 -2.992037e-02 -5.489333e-02
#> Locus_573 -2.158748e-02 -1.809304e-02 -3.102234e-03 -1.689569e-02
#> Locus_579 -1.479615e-03 -5.441758e-02 -1.327712e-03 1.615004e-01
#> Locus_581 -3.345055e-02 -4.723137e-02 -2.383616e-02 -8.222878e-02
#> Locus_582 -2.158748e-02 -1.809304e-02 -3.102234e-03 -1.689569e-02
#> Locus_583 -5.702073e-03 -3.684035e-03 1.010462e-02 -1.083814e-01
#> Locus_586 -3.715628e-03 -3.838249e-02 2.531179e-02 -5.813201e-02
#> Locus_589 2.223519e-02 -5.152151e-02 -3.656233e-03 -1.053461e-01
#> Locus_596 -7.428952e-03 -2.434339e-02 9.871652e-06 1.268967e-01
#> Locus_597 6.424873e-02 -2.242753e-02 -4.586720e-03 -1.925393e-02
#> Locus_598 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_600 5.464589e-03 -9.783744e-03 3.111483e-02 8.297926e-02
#> Locus_602 -7.776343e-02 -2.551947e-02 3.584306e-02 -3.856010e-02
#> Locus_603 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_604 6.068743e-03 4.563182e-03 -2.774035e-03 -3.765834e-03
#> Locus_609 -9.006329e-02 -4.464079e-02 -1.005676e-02 -4.562350e-02
#> Locus_612 -1.417826e-02 1.566581e-01 -2.711046e-02 9.724494e-02
#> Locus_613 2.798803e-03 1.091183e-02 5.931481e-02 -2.540233e-02
#> Locus_614 2.160314e-03 -1.942303e-03 9.348359e-03 1.298091e-01
#> Locus_622 -2.662558e-02 -1.472078e-02 -7.211615e-03 -3.728621e-03
#> Locus_625 -3.052196e-03 2.208376e-02 2.688017e-02 5.629993e-03
#> Locus_626 -1.108845e-01 -5.666179e-02 -3.395697e-02 -2.092199e-02
#> Locus_633 7.736958e-04 -7.750988e-03 -8.343130e-03 3.225658e-02
#> Locus_634 2.784287e-02 -2.145073e-03 1.008996e-02 -1.267557e-02
#> Locus_635 -8.376453e-02 -1.448553e-01 5.992058e-02 -1.719057e-02
#> Locus_636 2.123642e-01 -3.404766e-01 -5.060718e-03 7.424777e-02
#> Locus_637 5.257442e-02 7.160840e-03 -3.915763e-03 -2.514487e-02
#> Locus_639 -2.793866e-03 -2.998469e-02 -8.898187e-03 1.291245e-01
#> Locus_642 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_647 -4.975454e-02 -2.399596e-02 1.234110e-02 -1.826553e-02
#> Locus_652 1.368839e-02 -1.510601e-02 1.406611e-02 8.679250e-02
#> Locus_665 1.126052e-02 -1.234392e-02 3.651064e-04 -4.558033e-03
#> Locus_671 -8.848468e-02 -4.524291e-02 -1.978618e-02 -1.320049e-02
#> Locus_679 2.284531e-02 2.405548e-02 -3.509302e-02 -2.152929e-02
#> Locus_681 -1.352101e-02 2.038963e-02 3.509504e-02 5.238945e-01
#> Locus_682 -8.848468e-02 -4.524291e-02 -1.978618e-02 -1.320049e-02
#> Locus_683 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_685 -2.158748e-02 -1.809304e-02 -3.102234e-03 -1.689569e-02
#> Locus_687 5.080189e-03 2.659529e-02 -5.380601e-03 3.487952e-02
#> Locus_691 1.165835e-02 6.379972e-02 1.139152e-02 6.126231e-02
#> Locus_699 2.751067e-05 -1.126174e-01 -1.102065e-01 -3.731070e-02
#> Locus_701 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_707 6.075034e-02 -6.111350e-02 -2.885166e-02 -4.949400e-02
#> Locus_710 -1.697325e-03 -3.293379e-02 -5.867697e-02 -1.636054e-02
#> Locus_711 -3.569499e-02 -2.236516e-03 2.985183e-02 -1.663829e-02
#> Locus_712 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_717 2.284531e-02 2.405548e-02 -3.509302e-02 -2.152929e-02
#> Locus_718 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_724 1.326908e-01 -9.238781e-02 2.550381e-02 1.484621e-02
#> Locus_725 2.784287e-02 -2.145073e-03 1.008996e-02 -1.267557e-02
#> Locus_727 3.253208e-02 -2.722448e-02 -5.262506e-03 -1.266077e-02
#> Locus_729 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_733 -1.665339e-01 -6.602508e-02 8.211968e-03 -4.365386e-02
#> Locus_734 -5.468432e-03 -1.106160e-02 5.568668e-03 6.406873e-02
#> Locus_735 6.120651e-02 3.812663e-03 4.003859e-02 -2.504113e-02
#> Locus_736 -8.848468e-02 -4.524291e-02 -1.978618e-02 -1.320049e-02
#> Locus_740 -2.281511e-01 -6.426209e-02 -1.273478e-01 -2.423552e-01
#> Locus_742 -7.908979e-03 5.313441e-02 -2.098582e-02 -4.330562e-03
#> Locus_746 8.580060e-02 4.335044e-02 7.462346e-02 -8.571786e-02
#> Locus_750 -1.986935e-02 -1.482855e-02 5.189881e-03 -1.174941e-02
#> Locus_751 -5.325116e-02 -2.944157e-02 -1.442323e-02 -7.457241e-03
#> Locus_753 -6.916925e-03 5.275347e-02 -1.289708e-02 1.637812e-01
#> Locus_758 2.473155e-02 9.305913e-03 -1.400573e-02 -1.246929e-02
#> Locus_765 2.534445e-02 1.288367e-02 -1.202176e-02 -1.782733e-02
#> Locus_769 8.002374e-03 1.131640e-02 4.261605e-02 -2.700034e-02
#> Locus_770 1.629055e-01 -2.722181e-01 -1.440945e-02 1.994887e-01
#> Locus_771 -2.602901e-01 -1.759165e-01 -8.558347e-02 -4.312999e-02
#> Locus_776 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_779 -1.569730e-02 1.269421e-01 -2.958871e-02 1.381904e-02
#> Locus_784 -3.086843e-02 1.644903e-02 -2.262099e-02 -1.243771e-02
#> Locus_786 -1.985693e-01 -9.541836e-02 -1.087206e-03 -4.224222e-02
#> Locus_787 -3.417071e-02 -4.321145e-02 -2.260610e-02 -1.856373e-02
#> Locus_798 -4.354072e-02 1.086038e-01 1.483692e-02 5.491977e-01
#> Locus_802 8.878689e-03 5.802935e-02 1.645048e-02 -4.891675e-02
#> Locus_803 6.255389e-03 -2.023812e-02 6.987731e-03 -2.957126e-02
#> Locus_807 -3.973869e-02 -2.965711e-02 1.037976e-02 -2.349882e-02
#> Locus_808 3.193004e-02 2.291485e-02 -5.656485e-02 -1.534179e-02
#> Locus_809 -5.325116e-02 -2.944157e-02 -1.442323e-02 -7.457241e-03
#> Locus_811 -1.666692e-03 -3.340579e-02 -6.678782e-03 1.384450e-01
#> Locus_816 -4.657210e-03 4.242478e-02 2.418596e-03 1.195884e-02
#> Locus_821 -2.734216e-03 -5.530802e-03 2.784334e-03 3.057136e-02
#> Locus_825 5.962092e-02 4.915377e-02 -7.066688e-02 -5.321856e-02
#> Locus_831 -1.010649e-01 -7.740726e-02 1.765729e-02 -5.942573e-02
#> Locus_835 -2.385655e-02 -2.460049e-02 2.899625e-02 -2.288581e-02
#> Locus_838 -1.029442e-01 -4.539897e-02 -1.250874e-02 -2.385255e-02
#> Locus_847 -1.510607e-01 1.679202e-01 -1.354376e-02 3.579722e-02
#> Locus_850 -7.788322e-03 7.380767e-02 -8.602892e-03 2.141512e-02
#> Locus_851 5.665439e-02 -4.106422e-02 -8.803177e-03 -2.538164e-02
#> Locus_853 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_854 -1.059937e-02 3.079160e-02 7.280110e-03 -2.531519e-02
#> Locus_859 1.073239e-01 -7.623381e-02 3.729670e-02 -2.123165e-02
#> Locus_861 -9.393336e-03 9.414869e-02 -3.306446e-02 2.350585e-02
#> Locus_864 1.156102e-01 -1.149983e-01 2.723997e-02 -6.931959e-02
#> Locus_869 -5.200637e-02 -2.047813e-02 2.204672e-03 -7.407236e-03
#> Locus_870 -6.304430e-02 -5.101464e-02 -1.014587e-03 -4.554078e-02
#> Locus_875 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_876 -3.114321e-02 2.540998e-02 -2.303089e-02 -4.057898e-02
#> Locus_879 5.690695e-02 -8.954534e-02 -6.488892e-03 1.588098e-01
#> Locus_881 2.619006e-01 -1.219773e-03 -1.119716e-02 -4.149258e-01
#> Locus_887 4.450448e-02 -1.498322e-02 3.722497e-02 -2.259245e-02
#> Locus_888 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_895 1.355321e-02 -3.873580e-02 -3.692892e-02 -1.019406e-02
#> Locus_896 1.071830e-01 -9.724174e-02 2.351462e-03 -6.957475e-02
#> Locus_897 2.204501e-02 5.469886e-03 -7.456130e-04 2.129586e-02
#> Locus_898 7.729308e-03 -2.784120e-02 -2.115217e-02 -4.341403e-02
#> Locus_899 5.552053e-02 -1.872945e-02 2.103206e-02 -2.466826e-02
#> Locus_901 1.015908e-01 8.412897e-02 -9.281101e-02 -9.213068e-02
#> Locus_902 -1.614632e-01 -8.846402e-02 5.179238e-04 2.178047e-02
#> Locus_903 -7.364367e-02 -2.022304e-02 6.265524e-03 -7.085597e-02
#> Locus_905 -1.959494e-02 -2.367004e-02 1.463673e-02 1.434528e-01
#> Locus_908 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_909 5.150540e-02 -4.138249e-03 3.540968e-02 1.817243e-01
#> Locus_915 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_924 2.769258e-02 -2.405348e-02 -2.174791e-02 3.047912e-04
#> Locus_932 -2.893876e-02 -2.344287e-03 4.225332e-02 -2.465908e-02
#> Locus_935 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_936 -2.368680e-02 3.910875e-02 7.158028e-02 -4.670318e-02
#> Locus_938 5.568574e-02 -4.290147e-03 2.017993e-02 -2.535115e-02
#> Locus_941 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_944 -1.749158e-02 -9.002848e-03 -7.353265e-03 -1.577588e-02
#> Locus_945 -3.133488e-01 -8.640124e-02 -3.966238e-02 -9.501651e-02
#> Locus_946 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_949 -4.260900e-03 2.602541e-02 4.504992e-03 -2.213211e-02
#> Locus_950 -1.600230e-02 1.270661e-02 3.192763e-02 5.104617e-01
#> Locus_951 5.032894e-02 7.731251e-02 3.486011e-03 1.801274e-01
#> Locus_953 -7.516399e-03 3.343903e-02 -3.187232e-02 2.568046e-02
#> Locus_958 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_965 -8.019571e-03 -2.953769e-02 -1.159288e-02 1.296009e-01
#> Locus_966 5.790388e-02 -5.087152e-02 -2.445351e-03 -2.680372e-02
#> Locus_967 6.392933e-03 -3.515837e-02 -2.884926e-02 -1.126750e-02
#> Locus_968 2.550414e-02 3.143656e-02 4.977063e-02 -3.690078e-02
#> Locus_970 2.997505e-02 -2.784558e-02 -7.783785e-03 -3.586115e-02
#> Locus_972 2.742869e-03 -1.584921e-02 1.120955e-02 8.925329e-02
#> Locus_974 -4.145683e-02 -3.292160e-02 2.087647e-03 -2.864509e-02
#> Locus_979 -1.960520e-03 -1.328179e-02 -5.558796e-03 5.892071e-02
#> Locus_992 1.368839e-02 -1.510601e-02 1.406611e-02 8.530824e-02
#> Locus_998 -6.104392e-03 4.416752e-02 5.376034e-02 1.125999e-02
#> Locus_999 1.741142e-02 8.937418e-03 6.009702e-02 -2.796326e-02
#> Locus_1002 2.267980e-02 3.936505e-02 -2.685985e-02 -1.817463e-02
#> Locus_1005 1.476494e-01 -2.032025e-01 -7.340522e-02 -9.293945e-02
#> Locus_1006 -6.203602e-02 -3.242766e-02 4.009420e-02 2.310663e-02
#> Locus_1007 7.018085e-02 -5.441240e-02 7.007581e-02 -4.560841e-02
#> Locus_1015 3.983902e-02 -2.265757e-02 2.633039e-02 -3.001741e-02
#> Locus_1016 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1017 -9.892363e-02 -5.686227e-02 4.062280e-03 -3.112333e-02
#> Locus_1023 2.784287e-02 -2.145073e-03 1.008996e-02 -1.267557e-02
#> Locus_1024 -5.182862e-03 6.144204e-02 -2.387561e-02 1.805743e-03
#> Locus_1025 -4.682266e-02 2.505384e-02 4.403352e-02 8.034841e-02
#> Locus_1027 2.683097e-02 -1.905845e-02 9.324176e-03 -6.496173e-03
#> Locus_1030 1.160475e-01 -1.376418e-01 -1.959976e-02 4.780645e-01
#> Locus_1031 1.741133e-02 -4.418402e-02 -5.441605e-02 -2.255804e-02
#> Locus_1033 2.817218e-02 1.088358e-02 2.996862e-02 1.860167e-01
#> Locus_1034 -7.524399e-02 4.044818e-02 -5.103818e-02 -1.780477e-02
#> Locus_1035 2.252105e-02 -2.468785e-02 7.302129e-04 -9.524810e-03
#> Locus_1038 6.068743e-03 4.563182e-03 -2.774035e-03 -3.765834e-03
#> Locus_1041 -8.205925e-03 3.332350e-02 6.391176e-02 1.204839e-01
#> Locus_1042 -1.168116e-02 2.510986e-02 -3.247093e-02 -1.310481e-02
#> Locus_1044 -1.469603e-03 8.319893e-02 -2.098636e-03 -6.710055e-02
#> Locus_1046 2.683097e-02 -1.905845e-02 9.324176e-03 -4.119649e-03
#> Locus_1051 5.366195e-02 -3.811691e-02 1.864835e-02 -1.061582e-02
#> Locus_1054 1.248765e-02 4.517286e-03 -2.935880e-02 2.471051e-02
#> Locus_1055 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1065 9.256403e-03 -3.310299e-02 -2.688910e-02 -3.216045e-02
#> Locus_1071 4.946310e-02 1.861183e-02 -2.801145e-02 -2.493859e-02
#> Locus_1072 9.954069e-03 8.859504e-02 2.674415e-02 1.434821e-01
#> Locus_1073 -1.041283e-01 -5.676956e-02 -2.155547e-02 -2.670898e-02
#> Locus_1074 4.958153e-02 -3.959315e-02 -3.358506e-03 -1.583430e-02
#> Locus_1075 -2.661076e-02 -1.922967e-02 3.689166e-02 1.400823e-01
#> Locus_1076 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1080 4.918613e-02 5.080395e-03 -2.323982e-02 -1.700273e-02
#> Locus_1082 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1093 1.841220e-03 -3.562598e-02 -1.780625e-02 1.372043e-01
#> Locus_1095 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1098 8.002374e-03 1.131640e-02 4.261605e-02 -2.700034e-02
#> Locus_1099 -1.494260e-02 -4.583701e-03 3.575609e-03 -3.258059e-03
#> Locus_1105 9.907230e-02 2.216692e-02 6.653734e-02 -4.572362e-03
#> Locus_1106 1.872737e-02 4.582494e-02 5.375234e-03 1.687901e-02
#> Locus_1107 7.628746e-03 9.119301e-03 3.779691e-03 6.574041e-02
#> Locus_1108 -4.317496e-02 -3.618608e-02 -6.204467e-03 -3.379137e-02
#> Locus_1109 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1113 1.204341e-02 3.347834e-04 1.756686e-02 2.054127e-01
#> Locus_1115 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1118 -2.988520e-02 -9.167402e-03 7.151217e-03 -1.051993e-02
#> Locus_1122 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1127 -2.158748e-02 -1.809304e-02 -3.102234e-03 -1.689569e-02
#> Locus_1138 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1141 -5.504193e-02 -3.969824e-02 4.160146e-02 -7.030733e-02
#> Locus_1143 2.865791e-02 -7.402421e-02 -6.059539e-02 -3.689675e-02
#> Locus_1144 3.605672e-04 -3.653475e-02 -3.058819e-02 1.459312e-01
#> Locus_1155 2.320545e-02 2.034779e-02 -5.656433e-04 -8.021539e-03
#> Locus_1158 -1.248994e-02 1.535588e-01 -4.338532e-02 2.777872e-02
#> Locus_1159 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1161 3.663233e-02 -3.599096e-02 3.182261e-03 -1.829224e-02
#> Locus_1162 1.670953e-02 -6.635142e-02 -7.293598e-02 -2.962582e-02
#> Locus_1166 2.830997e-02 -2.471260e-02 2.269107e-03 -7.731570e-03
#> Locus_1172 -3.020119e-03 -2.134480e-02 -9.535754e-04 1.489222e-01
#> Locus_1179 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1181 2.683097e-02 -1.905845e-02 9.324176e-03 -4.119649e-03
#> Locus_1186 8.956451e-02 2.483285e-02 -2.977944e-02 -4.812881e-02
#> Locus_1191 -1.526098e-03 1.104188e-02 1.344008e-02 4.447754e-03
#> Locus_1197 -1.811429e-02 -2.276127e-02 2.741867e-02 1.249666e-01
#> Locus_1199 1.780922e-02 -3.766521e-03 2.897326e-02 1.566997e-01
#> Locus_1201 3.031339e-02 1.006541e-01 -7.287027e-02 -8.021542e-02
#> Locus_1204 1.547392e-03 -1.550198e-02 -1.668626e-02 5.767993e-02
#> Locus_1206 -3.086843e-02 1.644903e-02 -2.262099e-02 -1.243771e-02
#> Locus_1211 -5.624598e-02 -2.115486e-02 1.896926e-03 -5.514983e-03
#> Locus_1212 7.106981e-03 1.488864e-02 -1.452991e-03 9.862435e-02
#> Locus_1217 -4.117337e-02 -1.172230e-02 8.109391e-02 7.898646e-02
#> Locus_1223 1.561097e-02 2.256848e-02 1.812192e-02 1.026928e-01
#> Locus_1224 3.844744e-02 -1.910979e-02 3.075047e-02 -1.816124e-02
#> Locus_1227 -5.189593e-02 5.010932e-02 5.205277e-03 5.990920e-02
#> Locus_1231 -4.317496e-02 -3.618608e-02 -6.204467e-03 -3.379137e-02
#> Locus_1235 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1236 6.564431e-02 7.916838e-02 -9.415611e-02 -6.736952e-02
#> Locus_1239 -3.958559e-02 -2.809201e-02 5.563543e-02 2.343234e-02
#> Locus_1247 -2.063866e-01 -1.273514e-01 -5.145601e-02 1.051415e-01
#> Locus_1248 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1249 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1250 -6.801828e-03 2.685142e-02 7.176130e-03 1.434243e-01
#> Locus_1254 -4.145683e-02 -3.292160e-02 2.087647e-03 -2.864509e-02
#> Locus_1255 1.398485e-01 -4.818862e-02 7.616469e-02 -6.070704e-02
#> Locus_1256 3.277289e-02 5.460872e-02 1.418099e-02 2.551349e-02
#> Locus_1258 4.210454e-02 -3.669296e-02 -8.846113e-03 -9.583052e-02
#> Locus_1262 2.765161e-02 -4.378225e-02 -1.063676e-02 -3.198147e-02
#> Locus_1263 -1.984410e-02 3.013938e-01 3.083602e-02 1.230729e-02
#> Locus_1265 5.464589e-03 -9.783744e-03 3.111483e-02 8.223713e-02
#> Locus_1266 -3.722495e-02 1.607082e-02 6.849502e-05 -2.904381e-02
#> Locus_1270 4.564883e-03 2.620367e-02 -4.689373e-03 -2.089268e-02
#> Locus_1278 -2.965867e-01 -1.305725e-01 -1.521022e-02 -6.384104e-02
#> Locus_1279 -1.124361e-01 -5.664673e-02 -2.074071e-02 -2.613737e-02
#> Locus_1284 -9.663208e-02 -3.753812e-02 1.791301e-02 -2.335501e-02
#> Locus_1286 9.203228e-03 -3.544429e-02 -4.929207e-02 -6.172439e-02
#> Locus_1287 3.802680e-02 6.455464e-02 5.635741e-04 -2.201703e-01
#> Locus_1290 2.784287e-02 -2.145073e-03 1.008996e-02 -1.267557e-02
#> Locus_1295 8.211301e-03 -4.787603e-03 5.640887e-03 2.811057e-02
#> Locus_1301 -1.655891e-02 -1.489738e-02 -2.875915e-02 5.349155e-02
#> Locus_1302 1.700499e-02 3.295290e-02 -3.089590e-02 4.811064e-04
#> Locus_1305 7.108182e-03 7.527435e-02 1.611897e-02 7.865455e-02
#> Locus_1313 -1.008932e-02 3.221483e-02 -4.391634e-03 1.014664e-02
#> Locus_1315 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1316 -5.166936e-02 -1.415227e-02 -2.169257e-02 -1.667702e-01
#> Locus_1318 -3.115293e-01 -1.351562e-01 -1.163461e-02 -6.709910e-02
#> Locus_1319 -8.291365e-02 -6.584319e-02 4.175294e-03 -5.729019e-02
#> Locus_1338 -6.845691e-02 -6.727757e-02 2.166105e-02 -4.156941e-02
#> Locus_1343 -2.436133e-02 -5.695777e-02 5.345483e-02 -8.379319e-02
#> Locus_1345 1.285504e-02 -3.180890e-02 1.072672e-02 1.516048e-01
#> Locus_1347 -1.179136e-02 -1.691144e-02 1.649367e-02 9.305465e-02
#> Locus_1353 -3.234266e-03 4.908512e-02 7.731187e-03 4.363046e-02
#> Locus_1354 3.831311e-02 -1.887155e-02 3.131931e-02 -2.414129e-02
#> Locus_1355 -1.646870e-02 6.458179e-03 1.701569e-02 1.189696e-03
#> Locus_1360 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1363 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1364 -4.115575e-03 -3.396345e-03 8.715895e-03 5.524888e-02
#> Locus_1366 -2.297130e-02 9.483069e-02 -1.094998e-04 1.286393e-01
#> Locus_1370 1.353495e-01 7.499108e-02 -1.089949e-01 -1.420814e-01
#> Locus_1374 -2.274266e-01 -1.362364e-01 2.868382e-02 -7.117501e-02
#> Locus_1376 -2.906368e-02 -4.748920e-02 -3.752419e-03 -4.296755e-02
#> Locus_1377 2.451707e-02 1.673219e-02 -2.371591e-02 -2.110196e-02
#> Locus_1379 -9.413032e-03 8.113690e-02 -4.154602e-02 7.415380e-02
#> Locus_1380 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1382 6.569365e-02 2.779644e-02 1.670683e-02 -5.546186e-02
#> Locus_1384 -9.924487e-02 1.299396e-02 6.014516e-02 -5.231821e-02
#> Locus_1392 9.568341e-02 -1.126002e-01 1.332303e-02 -6.449065e-02
#> Locus_1396 1.244810e-02 1.307227e-01 -1.853443e-02 -1.134450e-01
#> Locus_1399 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1404 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1408 1.363411e-02 -1.102400e-02 7.193624e-02 3.324558e-01
#> Locus_1414 8.130716e-02 2.212495e-02 3.386534e-03 -5.111433e-02
#> Locus_1416 -1.705500e-02 -4.393913e-02 -6.618151e-02 -8.264783e-03
#> Locus_1420 1.395247e-02 -1.411095e-02 -5.407924e-03 -4.957967e-02
#> Locus_1423 -1.028921e-01 1.281987e-02 2.130370e-02 8.385242e-02
#> Locus_1424 2.085196e-02 1.602296e-01 -5.834472e-02 -3.417156e-02
#> Locus_1425 1.909717e-02 -2.849829e-02 -3.185081e-03 1.227186e-01
#> Locus_1426 3.382258e-02 -1.412110e-01 2.675842e-03 -1.384848e-01
#> Locus_1427 -2.662558e-02 -1.472078e-02 -7.211615e-03 -3.728621e-03
#> Locus_1430 -1.893140e-02 -1.426119e-01 -5.083465e-02 -3.325457e-01
#> Locus_1433 -1.498567e-02 5.739705e-02 3.326376e-02 -8.642124e-02
#> Locus_1439 -2.811896e-02 2.106761e-01 -6.166719e-02 4.373750e-03
#> Locus_1447 9.470409e-02 -4.198610e-02 9.047420e-02 1.188930e-01
#> Locus_1452 -1.811981e-02 2.368217e-02 2.085159e-02 6.017959e-02
#> Locus_1455 3.122359e-01 -1.065259e-01 1.948817e-02 -5.096994e-01
#> Locus_1457 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1459 -6.132617e-02 -4.775015e-02 7.277528e-03 -3.592691e-02
#> Locus_1460 -4.926790e-03 1.090151e-01 1.053879e-02 4.797524e-01
#> Locus_1461 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1463 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1468 -1.177483e-01 5.340747e-02 2.285353e-02 4.309195e-01
#> Locus_1470 -9.337443e-03 -4.421740e-02 -2.260681e-02 7.387953e-02
#> Locus_1477 2.010429e-01 -7.165281e-03 -6.589596e-02 -5.377491e-01
#> Locus_1479 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1482 7.736958e-04 -7.750988e-03 -8.343130e-03 2.542334e-02
#> Locus_1485 -7.788322e-03 7.380767e-02 -8.602892e-03 2.141512e-02
#> Locus_1486 2.463390e-02 -1.436281e-02 1.692266e-02 8.284744e-02
#> Locus_1487 1.137351e-02 4.433470e-02 2.947186e-02 2.109521e-01
#> Locus_1488 -4.606422e-02 -2.228689e-02 1.909293e-02 -1.548399e-02
#> Locus_1489 -8.333460e-04 -1.670290e-02 -3.339391e-03 6.629651e-02
#> Locus_1490 -6.585444e-04 -3.085767e-02 4.582940e-02 3.104543e-01
#> Locus_1494 1.174958e-02 2.045879e-02 1.868685e-02 1.297956e-01
#> Locus_1496 -2.110205e-02 1.741758e-01 6.681428e-02 1.559339e-01
#> Locus_1500 -9.393336e-03 9.414869e-02 -3.306446e-02 2.447845e-02
#> Locus_1502 -2.152272e-02 4.204910e-02 -2.026849e-03 -1.788931e-02
#> Locus_1506 -3.921040e-03 -2.656358e-02 -1.111759e-02 1.119894e-01
#> Locus_1508 -1.705500e-02 -4.393913e-02 -6.618151e-02 -8.264783e-03
#> Locus_1509 -9.927531e-03 2.018093e-02 7.976778e-02 1.549092e-03
#> Locus_1514 2.742869e-03 -1.584921e-02 1.120955e-02 9.143717e-02
#> Locus_1515 1.719630e-02 -1.732619e-02 2.938644e-03 8.164448e-02
#> Locus_1517 2.572774e-02 -1.135445e-02 2.387834e-02 -2.084124e-02
#> Locus_1520 -9.051978e-03 1.528962e-02 -9.406150e-03 3.919797e-02
#> Locus_1523 2.705259e-02 -6.527629e-03 3.095420e-02 -1.958326e-02
#> Locus_1524 -2.458508e-01 -7.324684e-02 1.091700e-02 4.931455e-01
#> Locus_1529 2.284531e-02 2.405548e-02 -3.509302e-02 -2.152929e-02
#> Locus_1534 7.455667e-03 4.707228e-02 -3.425656e-02 -3.503553e-03
#> Locus_1536 -1.417727e-01 5.526358e-04 7.105668e-02 3.385684e-02
#> Locus_1540 6.751028e-02 -2.757399e-02 6.910796e-02 -4.855604e-02
#> Locus_1545 -1.032351e-01 4.169228e-02 1.150926e-02 -1.219265e-01
#> Locus_1549 2.252105e-02 -2.468785e-02 7.302129e-04 -9.116066e-03
#> Locus_1551 -6.262224e-03 6.276579e-02 -2.204298e-02 1.502217e-02
#> Locus_1554 3.571938e-02 5.886785e-02 -2.432005e-02 -6.965100e-02
#> Locus_1555 -5.182862e-03 6.144204e-02 -2.387561e-02 1.805743e-03
#> Locus_1561 1.586801e-02 -4.549584e-02 1.488704e-02 3.124152e-02
#> Locus_1566 -9.328944e-03 -9.726945e-03 -1.496019e-02 -2.741034e-02
#> Locus_1572 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1573 -4.113747e-02 -1.204203e-02 1.747866e-02 -1.323025e-02
#> Locus_1576 1.029105e-02 -5.906496e-02 -3.872428e-02 -2.502389e-02
#> Locus_1586 -9.360142e-02 -2.592099e-02 1.620533e-02 -2.448951e-02
#> Locus_1587 2.674566e-03 -1.892308e-02 -1.446686e-02 6.798173e-02
#> Locus_1590 -4.424234e-02 -2.262146e-02 -9.893090e-03 -6.600247e-03
#> Locus_1591 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1596 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1597 2.365313e-01 2.161972e-02 -1.362514e-01 3.479165e-02
#> Locus_1598 -1.666692e-03 -3.340579e-02 -6.678782e-03 1.384450e-01
#> Locus_1601 9.210938e-02 -5.722670e-02 4.939882e-02 -2.877706e-02
#> Locus_1602 -1.526098e-03 1.104188e-02 1.344008e-02 4.447754e-03
#> Locus_1603 1.547392e-03 -1.550198e-02 -1.668626e-02 5.767993e-02
#> Locus_1604 2.640266e-02 6.419356e-02 -3.348393e-02 -3.318303e-02
#> Locus_1611 2.635265e-02 1.373990e-02 3.160182e-02 -4.594384e-02
#> Locus_1612 -4.777867e-03 2.175151e-02 -9.964330e-03 -1.184164e-02
#> Locus_1614 -6.636351e-02 -3.393219e-02 -1.483964e-02 -1.230938e-02
#> Locus_1618 -3.567562e-03 -2.223370e-02 -5.550573e-04 1.027199e-01
#> Locus_1619 9.047126e-02 -1.469682e-01 -3.193288e-02 3.618404e-02
#> Locus_1620 1.804091e-02 6.383583e-02 1.085782e-02 -7.198310e-02
#> Locus_1621 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1624 1.448380e-02 2.598959e-02 1.590251e-02 9.922423e-02
#> Locus_1626 -1.526098e-03 1.104188e-02 1.344008e-02 1.182239e-03
#> Locus_1632 7.480669e-03 3.011761e-02 -2.182228e-02 -2.235630e-02
#> Locus_1633 5.857690e-02 -5.759244e-02 6.763865e-02 3.640427e-02
#> Locus_1635 3.640635e-02 1.591881e-02 1.447293e-02 -6.957086e-03
#> Locus_1636 -6.715519e-02 3.627741e-02 -1.589202e-02 4.237239e-02
#> Locus_1641 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1644 -1.506232e-01 -8.631890e-02 -2.357721e-02 -3.995321e-02
#> Locus_1645 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1646 -1.219605e-01 -4.613028e-02 9.916464e-03 -2.736986e-02
#> Locus_1647 4.889964e-03 6.151521e-02 -1.152950e-02 1.090817e-01
#> Locus_1648 -1.142656e-02 4.928251e-02 3.389933e-02 2.321400e-01
#> Locus_1649 3.584524e-02 9.171325e-03 5.270601e-02 -3.967592e-02
#> Locus_1652 3.075673e-02 -5.995996e-02 -2.035418e-02 2.449764e-01
#> Locus_1656 2.473155e-02 9.305913e-03 -1.400573e-02 -1.246929e-02
#> Locus_1658 -2.559165e-02 -4.092521e-03 -2.140969e-02 3.519251e-03
#> Locus_1660 -1.710274e-02 1.586572e-01 -3.765701e-03 4.206728e-02
#> Locus_1661 4.542013e-02 7.947326e-03 2.870515e-02 -4.451115e-02
#> Locus_1664 5.471209e-02 -5.868756e-02 -1.472548e-02 -3.788222e-02
#> Locus_1665 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1670 1.011217e-02 -1.595970e-02 -4.828382e-04 6.383572e-02
#> Locus_1681 -3.217264e-03 3.873683e-02 -2.270378e-02 -2.050021e-02
#> Locus_1682 -1.084052e-02 9.589144e-02 1.827728e-02 2.509991e-02
#> Locus_1687 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1690 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1691 6.437033e-03 1.740619e-02 5.386488e-03 -2.398820e-02
#> Locus_1692 2.127155e-02 -1.488055e-02 -5.627613e-03 -8.102734e-03
#> Locus_1693 -2.159952e-02 6.453159e-02 1.716274e-02 -4.343937e-02
#> Locus_1694 -1.317609e-01 -5.900908e-02 -2.227561e-02 -2.658111e-02
#> Locus_1697 2.209948e-02 -3.511770e-02 2.349656e-02 3.028908e-01
#> Locus_1699 2.615936e-02 -5.044091e-03 -1.124235e-02 2.013861e-01
#> Locus_1702 8.652855e-06 -2.138001e-02 1.399389e-02 1.220085e-01
#> Locus_1703 4.797294e-03 4.311498e-02 -2.199788e-02 -1.380046e-02
#> Locus_1708 -4.239021e-03 1.050983e-03 1.297824e-02 9.031356e-02
#> Locus_1713 1.374122e-03 6.343345e-03 4.038110e-02 1.181818e-01
#> Locus_1715 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1724 -2.316749e-01 -1.705653e-03 3.017282e-02 -8.080255e-02
#> Locus_1728 -5.307839e-02 3.310132e-02 4.373521e-02 -6.494971e-02
#> Locus_1730 -2.189488e-02 -2.233175e-02 3.097040e-02 1.548956e-01
#> Locus_1734 -8.882395e-02 -2.604668e-02 1.258316e-02 -2.726554e-02
#> Locus_1737 -7.412754e-02 -3.178886e-02 -2.741873e-03 -8.298355e-03
#> Locus_1742 -1.185584e-01 -3.148584e-02 2.721666e-02 -3.575315e-02
#> Locus_1745 -8.291365e-02 -6.584319e-02 4.175294e-03 -5.729019e-02
#> Locus_1748 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1750 -6.555007e-02 -2.690687e-02 1.052731e-02 8.837082e-02
#> Locus_1751 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1755 1.547392e-03 -1.550198e-02 -1.668626e-02 6.451316e-02
#> Locus_1756 3.538283e-02 -2.618367e-02 -3.175564e-03 -1.727891e-02
#> Locus_1757 -1.059937e-02 3.079160e-02 7.280110e-03 -2.531519e-02
#> Locus_1774 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1776 8.652194e-02 -4.234074e-02 2.551445e-02 2.438030e-02
#> Locus_1777 2.742869e-03 -1.584921e-02 1.120955e-02 9.436318e-02
#> Locus_1778 -6.262224e-03 6.276579e-02 -2.204298e-02 1.599476e-02
#> Locus_1783 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1784 6.940446e-02 1.329712e-01 -5.241321e-02 -1.244828e-01
#> Locus_1786 -2.310285e-02 -2.144986e-02 2.576419e-02 1.349343e-01
#> Locus_1787 4.824879e-02 -3.604230e-02 2.460855e-02 -2.995730e-02
#> Locus_1797 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1799 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1801 -2.374018e-01 -1.033673e-01 -8.892737e-03 -5.880074e-02
#> Locus_1804 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1805 2.787562e-02 5.187846e-04 -3.111369e-02 -4.183427e-02
#> Locus_1807 -5.935835e-03 -3.576336e-02 4.119374e-02 4.278169e-01
#> Locus_1808 8.160859e-02 -1.565283e-01 3.792738e-02 1.179286e-02
#> Locus_1811 4.765276e-03 2.472947e-02 -5.990631e-03 -2.448818e-02
#> Locus_1813 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1816 2.572774e-02 -1.135445e-02 2.387834e-02 -2.084124e-02
#> Locus_1822 -7.234611e-03 9.150118e-02 -3.672973e-02 -2.927000e-03
#> Locus_1823 4.258452e-02 6.253918e-02 -6.877326e-02 -5.454555e-02
#> Locus_1826 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1828 4.457619e-02 7.122796e-03 9.522803e-03 -2.765729e-02
#> Locus_1833 -2.559522e-01 -1.225297e-02 -5.660365e-03 -9.161514e-02
#> Locus_1839 7.639327e-03 -1.284991e-02 -2.379676e-02 -1.018708e-02
#> Locus_1840 -3.673793e-02 -1.877990e-02 -8.179793e-04 -5.777957e-02
#> Locus_1841 1.158758e-01 -8.335903e-02 2.479696e-02 -3.201438e-02
#> Locus_1842 8.041729e-02 5.015767e-03 6.174201e-03 -3.782044e-02
#> Locus_1843 1.832347e-02 -2.074730e-02 5.158049e-03 9.046203e-02
#> Locus_1844 -1.515425e-01 -7.174492e-02 5.858972e-02 2.742252e-01
#> Locus_1846 -2.881668e-02 -1.361011e-02 -9.766868e-03 -6.732373e-03
#> Locus_1848 -7.709406e-03 6.450854e-02 2.929876e-02 1.758883e-02
#> Locus_1849 -3.676317e-02 -3.955723e-02 9.173633e-03 -4.198175e-02
#> Locus_1852 -4.799002e-02 1.619411e-01 7.288537e-02 4.935796e-01
#> Locus_1853 4.559642e-02 2.792469e-02 1.578829e-02 -1.520451e-02
#> Locus_1856 1.011217e-02 -1.595970e-02 -4.828382e-04 6.383572e-02
#> Locus_1861 7.377955e-03 -2.149050e-02 2.301496e-03 9.366495e-02
#> Locus_1863 -1.494260e-02 -4.583701e-03 3.575609e-03 -7.261874e-03
#> Locus_1868 -9.298985e-02 -5.909868e-02 -4.043469e-03 -2.648847e-02
#> Locus_1870 -1.526098e-03 1.104188e-02 1.344008e-02 4.447754e-03
#> Locus_1871 -4.121026e-03 -6.578102e-04 1.242197e-02 2.584711e-02
#> Locus_1872 -1.052575e-01 -4.991970e-02 -1.221856e-02 -1.968249e-02
#> Locus_1873 2.059894e-02 -6.129826e-02 5.199268e-02 5.463895e-01
#> Locus_1881 1.775796e-01 -2.197513e-01 -4.698121e-02 -3.028696e-01
#> Locus_1882 -2.619487e-02 -7.458334e-03 1.390305e-02 -5.968374e-03
#> Locus_1887 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1894 3.799405e-02 1.179817e-01 -7.943912e-02 -3.707821e-02
#> Locus_1896 1.547392e-03 -1.550198e-02 -1.668626e-02 5.767993e-02
#> Locus_1908 -4.209466e-02 6.401259e-02 3.088285e-02 7.380954e-02
#> Locus_1911 -5.977040e-02 -1.833480e-02 1.430243e-02 -2.103986e-02
#> Locus_1913 -5.493424e-03 -9.992283e-03 5.094789e-02 1.097374e-01
#> Locus_1919 -4.777867e-03 2.175151e-02 -9.964330e-03 -1.184164e-02
#> Locus_1921 2.327765e-02 7.295083e-03 2.340460e-02 8.970496e-02
#> Locus_1932 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1933 1.433771e-03 5.758656e-02 -1.571086e-02 -1.240900e-02
#> Locus_1935 -2.158748e-02 -1.809304e-02 -3.102234e-03 -1.689569e-02
#> Locus_1939 -2.746712e-03 -4.996142e-03 2.547395e-02 5.412657e-02
#> Locus_1941 -1.370473e-02 -5.204680e-03 4.530701e-02 8.088470e-02
#> Locus_1942 1.900870e-03 -1.117209e-02 -6.123725e-03 3.572515e-02
#> Locus_1945 2.202137e-02 6.895017e-03 2.733440e-02 -2.614912e-02
#> Locus_1948 2.451707e-02 1.673219e-02 -2.371591e-02 -2.110196e-02
#> Locus_1949 -1.687393e-01 -1.231910e-01 -4.891367e-02 -5.809020e-02
#> Locus_1950 -1.506098e-02 1.645321e-02 5.178894e-02 8.700009e-02
#> Locus_1951 -5.093660e-03 -1.119182e-02 1.288503e-02 9.805012e-02
#> Locus_1952 -9.038767e-03 4.777692e-02 -5.459339e-03 -3.397376e-02
#> Locus_1953 -1.813883e-02 2.496854e-02 7.412689e-02 -2.581935e-02
#> Locus_1958 4.221912e-02 -5.115581e-02 -1.359863e-02 -2.020371e-02
#> Locus_1961 -2.706329e-02 -3.273665e-02 3.113755e-02 2.205703e-01
#> Locus_1962 5.993070e-03 8.140283e-02 -1.465492e-02 -2.739945e-01
#> Locus_1965 4.774628e-03 -3.138699e-02 -3.242816e-02 -1.534963e-02
#> Locus_1967 -4.317496e-02 -3.618608e-02 -6.204467e-03 -3.379137e-02
#> Locus_1968 -1.370473e-02 -5.204680e-03 4.530701e-02 8.088470e-02
#> Locus_1971 4.234158e-02 -1.135290e-02 4.711351e-02 -2.068721e-02
#> Locus_1974 9.772001e-03 -3.133410e-02 -3.061928e-02 -1.237095e-02
#> Locus_1977 -8.313636e-02 -3.860897e-02 -7.272013e-03 -2.198099e-02
#> Locus_1982 5.089783e-02 9.663334e-03 -6.101867e-02 1.945626e-02
#> Locus_1983 1.577272e-02 2.001358e-02 -1.143140e-02 -6.201321e-02
#> Locus_1984 5.928265e-04 7.963608e-02 -9.920155e-03 4.947859e-02
#> Locus_1988 -3.973869e-02 -2.965711e-02 1.037976e-02 -2.126502e-02
#> Locus_1990 -1.048652e-02 1.406453e-02 5.294693e-02 6.038248e-02
#> Locus_1991 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1993 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2003 -6.444559e-03 -1.165428e-02 -1.664311e-02 1.207514e-01
#> Locus_2007 -4.317496e-02 -3.618608e-02 -6.204467e-03 -3.379137e-02
#> Locus_2014 8.049292e-02 -5.717536e-02 2.797253e-02 -1.473547e-02
#> Locus_2016 -1.108845e-01 -5.666179e-02 -3.395697e-02 -2.092199e-02
#> Locus_2017 4.923203e-02 -3.262460e-02 3.147271e-02 -3.832207e-02
#> Locus_2019 -1.489092e-02 2.152436e-02 5.430788e-02 -2.638322e-02
#> Locus_2020 1.499770e-02 1.769047e-02 -1.095012e-02 -8.470134e-03
#> Locus_2021 -1.942597e-02 1.213096e-02 -3.927184e-02 3.973181e-03
#> Locus_2027 -1.059937e-02 3.079160e-02 7.280110e-03 -2.531519e-02
#> Locus_2030 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2033 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2036 -4.657210e-03 4.242478e-02 2.418596e-03 8.693322e-03
#> Locus_2041 -7.788322e-03 7.380767e-02 -8.602892e-03 2.141512e-02
#> Locus_2045 -3.130554e-01 -1.241143e-01 1.805474e-03 -6.265134e-02
#> Locus_2048 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2050 5.568574e-02 -4.290147e-03 2.017993e-02 -2.535115e-02
#> Locus_2053 -1.277801e-01 1.584879e-01 8.035684e-02 -1.190989e-01
#> Locus_2055 -1.422099e-01 -7.943124e-02 -7.865841e-03 -4.067207e-02
#> Locus_2060 4.268653e-02 1.445094e-02 -6.665956e-02 -8.654309e-03
#> Locus_2063 -1.986935e-02 -1.482855e-02 5.189881e-03 -9.515613e-03
#> Locus_2064 9.716236e-02 -9.603220e-03 5.133997e-02 -4.505692e-02
#> Locus_2068 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2069 -9.314384e-02 -3.252017e-02 1.968333e-02 -2.545549e-02
#> Locus_2080 -6.272990e-02 -4.492370e-02 -2.716513e-02 -1.479207e-02
#> Locus_2085 7.628746e-03 9.119301e-03 3.779691e-03 6.574041e-02
#> Locus_2086 4.946310e-02 1.861183e-02 -2.801145e-02 -2.493859e-02
#> Locus_2087 2.473155e-02 9.305913e-03 -1.400573e-02 -1.246929e-02
#> Locus_2091 -4.716860e-03 1.797089e-02 -9.263925e-03 1.101724e-01
#> Locus_2094 -7.008327e-03 -1.199269e-02 3.342514e-03 8.453699e-02
#> Locus_2102 -4.338308e-02 -8.378164e-02 3.231059e-03 -3.464460e-02
#> Locus_2106 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2108 -5.977040e-02 -1.833480e-02 1.430243e-02 -2.103986e-02
#> Locus_2109 -1.494260e-02 -4.583701e-03 3.575609e-03 -3.258059e-03
#> Locus_2113 1.139428e-02 -6.676895e-02 -5.327844e-02 -1.067882e-02
#> Locus_2115 -1.960520e-03 -1.328179e-02 -5.558796e-03 6.282795e-02
#> Locus_2120 8.238213e-02 -5.241868e-02 1.507516e-02 -4.622288e-02
#> Locus_2128 1.775355e-02 3.006976e-02 5.698330e-03 -2.528935e-03
#> Locus_2131 7.677539e-03 9.769077e-02 4.407224e-02 4.058370e-01
#> Locus_2133 -1.027830e-01 -8.067175e-02 9.365174e-03 -6.457201e-02
#> Locus_2135 7.736958e-04 -7.750988e-03 -8.343130e-03 2.542334e-02
#> Locus_2136 -1.208126e-02 3.927019e-02 4.304523e-02 3.999565e-01
#> Locus_2137 -8.227494e-02 -2.408407e-02 3.495732e-02 -2.646050e-02
#> Locus_2138 6.437033e-03 1.740619e-02 5.386488e-03 -2.398820e-02
#> Locus_2139 -1.986935e-02 -1.482855e-02 5.189881e-03 -1.174941e-02
#> Locus_2140 -6.063483e-03 3.379960e-03 2.639709e-02 1.259959e-01
#> Locus_2145 -4.393565e-02 -6.226486e-02 3.246337e-03 -5.173827e-02
#> Locus_2148 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2154 -1.537724e-02 5.254311e-02 -2.684220e-03 -3.715684e-02
#> Locus_2157 -2.036863e-02 -1.591905e-02 2.297986e-02 1.180294e-01
#> Locus_2159 5.232859e-02 -3.380357e-02 1.097390e-02 -2.157116e-02
#> Locus_2162 7.014923e-02 -1.126971e-01 -1.631208e-02 -1.436799e-01
#> Locus_2165 2.321087e-03 -2.325296e-02 -2.502939e-02 8.310327e-02
#> Locus_2166 -2.051750e-03 3.005914e-02 -1.285412e-02 -5.705340e-03
#> Locus_2168 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2179 -2.838339e-02 5.145507e-02 -2.698952e-02 1.125823e-02
#> Locus_2180 1.173746e-01 -9.925591e-02 4.179804e-03 -4.606751e-02
#> Locus_2181 6.827085e-02 -4.111556e-02 1.262311e-02 -3.704671e-02
#> Locus_2183 6.491782e-02 -1.144441e-02 -1.775935e-03 -8.741710e-02
#> Locus_2185 -1.545997e-02 -4.069982e-02 -4.348373e-02 -6.290378e-02
#> Locus_2186 3.663233e-02 -3.599096e-02 3.182261e-03 -1.910973e-02
#> Locus_2188 6.855050e-03 1.687029e-02 1.212282e-02 3.348383e-02
#> Locus_2191 4.120834e-03 1.133949e-02 1.490716e-02 6.698120e-02
#> Locus_2203 4.527201e-02 -3.497491e-02 6.808097e-03 -2.111714e-02
#> Locus_2208 -1.779872e-02 9.672337e-02 2.490713e-02 2.773547e-02
#> Locus_2211 -3.821434e-02 1.382853e-01 3.253046e-02 2.585192e-01
#> Locus_2213 5.026938e-02 -3.492202e-02 8.616973e-03 -1.813846e-02
#> Locus_2215 9.399461e-03 -2.902524e-02 -1.012023e-02 -5.893624e-02
#> Locus_2216 2.596370e-02 5.620064e-03 1.638375e-02 2.728633e-02
#> Locus_2219 9.264113e-03 -8.347334e-03 -6.411907e-02 -9.200866e-02
#> Locus_2223 1.506635e-02 1.208269e-02 1.776371e-02 6.085226e-02
#> Locus_2228 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2229 1.225854e-02 8.366097e-03 -1.185795e-02 -1.051465e-02
#> Locus_2230 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2231 -2.734216e-03 -5.530802e-03 2.784334e-03 3.057136e-02
#> Locus_2234 8.211301e-03 -4.787603e-03 5.640887e-03 2.736844e-02
#> Locus_2236 1.535205e-02 4.041885e-02 -2.922570e-02 -3.550282e-02
#> Locus_2238 1.065679e-02 -5.473901e-03 -1.246288e-04 1.049341e-01
#> Locus_2243 -1.196733e-02 5.964206e-02 -3.142382e-03 4.840384e-02
#> Locus_2246 5.746548e-02 2.992822e-02 1.460460e-02 -5.193893e-02
#> Locus_2250 1.292413e-02 1.185114e-01 8.922477e-04 2.279484e-01
#> Locus_2251 -5.798908e-03 1.708762e-02 5.235412e-02 6.049869e-02
#> Locus_2254 4.968576e-02 -6.267254e-02 -1.384445e-02 -1.110464e-01
#> Locus_2260 -2.804152e-03 3.335003e-02 -7.757169e-03 2.090024e-02
#> Locus_2261 5.758661e-02 -1.546224e-02 1.405620e-02 1.037400e-02
#> Locus_2267 9.705730e-03 4.019247e-02 -1.473835e-02 -1.542349e-01
#> Locus_2268 -3.236970e-03 7.148898e-02 -3.391429e-02 2.870563e-01
#> Locus_2271 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2273 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2274 -6.418344e-02 -7.374201e-02 -7.476078e-02 -8.802391e-03
#> Locus_2275 5.435616e-02 -7.098015e-02 -3.578666e-02 -3.077519e-02
#> Locus_2277 4.775000e-02 1.662952e-02 1.913668e-02 -4.435944e-02
#> Locus_2282 6.978332e-02 6.022225e-02 9.115819e-02 1.295179e-01
#> Locus_2287 5.686745e-02 -8.694293e-02 4.972733e-02 -1.055439e-01
#> Locus_2294 2.787562e-02 5.187846e-04 -3.111369e-02 -4.183427e-02
#> Locus_2298 3.819664e-03 -6.424957e-03 -1.189838e-02 -5.093542e-03
#> Locus_2305 1.433864e-02 2.214563e-02 3.409192e-02 5.714291e-01
#> Locus_2308 -5.857338e-02 3.052001e-03 -2.799997e-02 -1.977313e-03
#> Locus_2309 -4.424234e-02 -2.262146e-02 -9.893090e-03 -6.600247e-03
#> Locus_2312 9.307171e-02 -8.593862e-02 -1.005867e-04 -6.481878e-02
#> Locus_2318 1.672657e-02 -8.097090e-02 -6.508620e-02 -2.234763e-02
#> Locus_2319 3.273918e-02 -2.642116e-02 6.436764e-02 -1.830517e-02
#> Locus_2321 8.246448e-02 -1.161098e-01 -9.890205e-02 -9.710539e-02
#> Locus_2322 -3.655215e-02 -8.450617e-02 -4.136895e-02 -7.222680e-02
#> Locus_2323 2.237444e-02 -1.320668e-02 1.565863e-02 5.431917e-02
#> Locus_2325 6.368811e-02 7.026252e-03 6.279598e-02 -5.235149e-02
#> Locus_2326 3.288802e-02 -1.493189e-02 1.579868e-02 -1.976780e-02
#> Locus_2327 1.651197e-02 -3.481803e-02 3.528349e-02 -7.264269e-02
#> Locus_2333 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2336 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2338 -4.329221e-01 3.865181e-02 -6.763400e-02 -1.295999e-01
#> Locus_2339 1.506635e-02 1.208269e-02 1.776371e-02 6.085226e-02
#> Locus_2344 -1.009059e-01 -6.748101e-04 -5.269822e-03 -1.087047e-02
#> Locus_2345 -2.619487e-02 -7.458334e-03 1.390305e-02 -5.968374e-03
#> Locus_2348 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2349 1.547392e-03 -1.550198e-02 -1.668626e-02 5.084669e-02
#> Locus_2354 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2359 6.685203e-03 6.254277e-03 1.908097e-02 3.181619e-02
#> Locus_2361 -7.540040e-02 -3.889398e-02 -2.432055e-02 -2.708074e-02
#> Locus_2364 2.845840e-02 2.430120e-02 3.272089e-02 -5.013732e-02
#> Locus_2365 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2366 -2.838128e-02 4.783726e-02 -4.302667e-04 1.178594e-01
#> Locus_2368 -8.352381e-02 3.220089e-02 2.369528e-02 5.224880e-02
#> Locus_2371 -6.262224e-03 6.276579e-02 -2.204298e-02 1.502217e-02
#> Locus_2375 4.752686e-03 7.121045e-02 -2.194559e-02 -6.081801e-02
#> Locus_2381 2.451707e-02 1.673219e-02 -2.371591e-02 -2.110196e-02
#> Locus_2388 -6.837715e-03 5.458748e-02 -2.292391e-02 1.916068e-02
#> Locus_2394 -4.026078e-02 -5.826080e-02 -7.286265e-02 -2.924260e-02
#> Locus_2400 1.177012e-01 -1.013649e-01 7.636215e-04 -5.096416e-02
#> Locus_2402 4.030795e-02 8.219104e-02 -4.639904e-02 -2.434496e-02
#> Locus_2406 7.702900e-02 2.935322e-03 -1.314985e-02 -2.967830e-02
#> Locus_2413 -4.375928e-02 -1.819381e-02 -6.191259e-03 -1.399425e-02
#> Locus_2414 -4.657210e-03 4.242478e-02 2.418596e-03 8.693322e-03
#> Locus_2415 -3.323198e-03 6.861094e-02 -3.207797e-02 -1.573996e-02
#> Locus_2421 -4.145683e-02 -3.292160e-02 2.087647e-03 -2.641130e-02
#> Locus_2424 -7.788322e-03 7.380767e-02 -8.602892e-03 1.946992e-02
#> Locus_2425 1.881240e-01 -1.489808e-01 7.439146e-03 -7.153201e-02
#> Locus_2426 -6.304430e-02 -5.101464e-02 -1.014587e-03 -4.554078e-02
#> Locus_2431 -2.506498e-01 -1.148588e-01 -2.796196e-02 -4.794025e-02
#> Locus_2432 -6.262224e-03 6.276579e-02 -2.204298e-02 1.599476e-02
#> Locus_2433 -8.283220e-02 -2.747401e-02 2.176058e-03 -9.567186e-03
#> Locus_2434 -4.317496e-02 -3.618608e-02 -6.204467e-03 -3.379137e-02
#> Locus_2435 -1.494260e-02 -4.583701e-03 3.575609e-03 -7.261874e-03
#> Locus_2436 -5.681425e-02 -9.670618e-02 -7.969595e-02 -4.300864e-02
#> Locus_2437 -5.977040e-02 -1.833480e-02 1.430243e-02 -2.103986e-02
#> Locus_2439 -8.608822e-03 -2.717248e-02 5.554987e-03 -1.407365e-02
#> Locus_2448 6.197417e-02 2.928468e-02 -8.508819e-02 -2.044101e-01
#> Locus_2449 1.001236e-02 -2.544749e-02 -8.136258e-03 -6.429427e-02
#> Locus_2450 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2451 -1.120396e-02 -9.794874e-04 1.998735e-03 2.890295e-01
#> Locus_2452 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2457 1.513764e-02 7.118133e-04 -7.867715e-02 -1.334150e-01
#> Locus_2461 -3.706377e-02 -1.589443e-02 -1.370936e-03 -4.149178e-03
#> Locus_2463 5.968692e-02 1.368053e-03 4.148795e-02 -3.885132e-02
#> Locus_2466 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2467 -1.696911e-01 -7.376969e-02 -1.746946e-03 -3.186962e-02
#> Locus_2473 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2475 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2477 -1.587523e-01 1.581996e-02 -4.853014e-02 1.590185e-01
#> Locus_2480 1.608449e-01 -1.353586e-01 2.099981e-02 -8.616982e-02
#> Locus_2494 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2497 2.346879e-02 1.345100e-02 1.320027e-02 1.588493e-01
#> Locus_2501 7.052940e-02 1.230586e-02 -5.656960e-02 -2.132988e-02
#> Locus_2502 3.801740e-03 -2.234419e-02 -1.224745e-02 7.145030e-02
#> Locus_2507 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2510 4.450448e-02 -1.498322e-02 3.722497e-02 -3.143287e-02
#> Locus_2512 -4.334680e-02 4.669112e-02 -7.343817e-03 3.870743e-01
#> Locus_2514 -2.542897e-03 7.710360e-02 -3.844769e-02 -2.006924e-02
#> Locus_2517 8.829708e-02 -5.455162e-02 3.232757e-02 -4.946915e-02
#> Locus_2518 -5.865328e-03 2.585210e-02 -8.237154e-03 3.808245e-02
#> Locus_2519 5.794916e-02 5.647705e-02 -8.204400e-02 -5.371854e-02
#> Locus_2522 8.530835e-02 2.778315e-02 2.469456e-02 -6.461450e-02
#> Locus_2526 -1.040127e-01 -4.095626e-02 4.409345e-03 -2.282210e-02
#> Locus_2529 7.480669e-03 3.011761e-02 -2.182228e-02 -2.242895e-02
#> Locus_2530 -1.244553e-02 1.162325e-01 -6.184297e-03 3.337395e-02
#> Locus_2532 6.055250e-02 -6.497081e-02 -1.867819e-02 -3.471782e-02
#> Locus_2540 -5.076038e-02 -7.693092e-03 -1.029930e-02 5.658953e-02
#> Locus_2542 -3.052196e-03 2.208376e-02 2.688017e-02 5.629993e-03
#> Locus_2546 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2550 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2555 -8.227494e-02 -2.408407e-02 3.495732e-02 -1.845287e-02
#> Locus_2559 -2.970617e-01 -5.258203e-02 1.260406e-01 1.190481e-02
#> Locus_2561 6.226459e-02 1.067906e-01 -6.636517e-02 -1.044741e-01
#> Locus_2565 2.784287e-02 -2.145073e-03 1.008996e-02 -1.267557e-02
#> Locus_2566 -4.317496e-02 -3.618608e-02 -6.204467e-03 -3.379137e-02
#> Locus_2567 -6.258781e-02 -4.952544e-02 -4.930344e-02 -1.363070e-02
#> Locus_2572 -8.848468e-02 -4.524291e-02 -1.978618e-02 -1.320049e-02
#> Locus_2573 -6.289673e-02 -6.095295e-02 1.098844e-02 3.362050e-01
#> Locus_2577 4.821018e-02 -3.604047e-02 6.260049e-03 -1.470576e-02
#> Locus_2579 2.767678e-02 8.555462e-02 3.407937e-02 -1.029073e-01
#> Locus_2581 1.538448e-02 -2.389028e-02 -1.740955e-02 -1.464934e-01
#> Locus_2583 1.975183e-02 -1.003254e-02 1.239744e-02 -1.913322e-02
#> Locus_2588 1.419590e-01 -9.266853e-02 5.097592e-02 -5.770845e-02
#> Locus_2589 -7.939374e-02 -2.038826e-02 1.532812e-02 -1.085192e-01
#> Locus_2592 -1.590332e-02 2.359126e-02 5.071330e-02 1.099803e-02
#> Locus_2597 -2.051750e-03 3.005914e-02 -1.285412e-02 -5.705340e-03
#> Locus_2598 3.514381e-02 -5.498114e-02 -2.780213e-02 -2.651390e-02
#> Locus_2599 2.473155e-02 9.305913e-03 -1.400573e-02 -1.246929e-02
#> Locus_2601 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2602 -1.355261e-01 -5.352564e-02 2.053409e-02 -3.391774e-02
#> Locus_2611 3.484939e-02 -1.540080e-04 6.427888e-02 -2.615478e-02
#> Locus_2616 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2617 -5.155980e-04 -1.768886e-03 3.730178e-03 -1.978950e-02
#> Locus_2619 2.742869e-03 -1.584921e-02 1.120955e-02 9.436318e-02
#> Locus_2620 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2629 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2632 -2.509660e-02 3.367655e-02 6.654720e-02 6.867288e-02
#> Locus_2634 -1.946459e-02 3.222479e-02 -2.880892e-02 1.442697e-01
#> Locus_2636 -3.228614e-03 -3.137805e-02 -8.092572e-03 -2.970030e-02
#> Locus_2639 -1.204379e-02 2.102857e-02 3.709140e-02 -1.975608e-02
#> Locus_2644 7.693472e-03 2.517304e-02 -1.639507e-02 2.162803e-01
#> Locus_2647 -2.150431e-02 4.899513e-02 6.743415e-02 4.798026e-01
#> Locus_2648 8.755920e-03 5.698194e-03 5.999096e-03 6.920898e-02
#> Locus_2652 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2655 3.910339e-02 -1.448900e-02 1.045507e-02 -1.723361e-02
#> Locus_2656 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2657 -1.543816e-01 4.642340e-02 -2.158922e-02 1.197363e-01
#> Locus_2660 -2.793866e-03 -2.998469e-02 -8.898187e-03 1.281432e-01
#> Locus_2662 8.903789e-02 1.569590e-02 -6.799105e-02 -5.495095e-01
#> Locus_2665 5.347992e-03 1.840783e-01 -1.060264e-01 -1.131804e-01
#> Locus_2668 -3.964664e-02 3.477010e-02 -2.624901e-02 1.637164e-01
#> Locus_2669 -6.797850e-02 -1.167038e-01 3.396920e-02 -9.549316e-02
#> Locus_2670 -7.891246e-02 -6.018499e-02 2.548332e-02 -6.855657e-02
#> Locus_2671 2.473155e-02 9.305913e-03 -1.400573e-02 -1.246929e-02
#> Locus_2675 1.022142e-01 -3.680119e-02 -4.639189e-02 -1.648324e-01
#> Locus_2677 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2685 6.197388e-02 -2.186440e-02 -1.403599e-02 -8.408111e-02
#> Locus_2687 1.642260e-02 -9.575206e-03 1.128177e-02 5.547900e-02
#> Locus_2696 -3.627212e-03 -4.668758e-02 -1.223758e-02 1.944398e-01
#> Locus_2698 1.233286e-01 -9.114305e-02 5.854128e-02 -6.722827e-02
#> Locus_2699 -6.304430e-02 -5.101464e-02 -1.014587e-03 -4.554078e-02
#> Locus_2702 1.329149e-02 2.180769e-02 2.602855e-04 6.224796e-02
#> Locus_2703 1.298957e-01 3.106199e-02 -4.808873e-02 -3.754366e-02
#> Locus_2705 -2.313143e-04 -3.219536e-02 6.900482e-02 3.588129e-03
#> Locus_2712 1.586290e-02 2.058683e-02 4.546106e-03 1.602580e-01
#> Locus_2718 -1.278054e-03 2.230815e-02 -2.119725e-02 2.655124e-02
#> Locus_2719 1.231964e-02 7.086544e-03 4.323766e-02 1.164631e-01
#> Locus_2724 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2725 -1.824754e-02 2.653868e-02 4.901400e-03 4.685743e-02
#> Locus_2727 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2729 9.139836e-03 6.927530e-02 -2.640013e-02 -5.833146e-02
#> Locus_2731 3.144072e-03 -8.787129e-03 -1.710796e-02 -2.936498e-02
#> Locus_2733 -1.812427e-01 -9.757911e-02 -7.084709e-03 -8.882993e-03
#> Locus_2736 -2.684064e-03 3.782396e-03 -8.282344e-03 -1.384537e-02
#> Locus_2738 -5.325116e-02 -2.944157e-02 -1.442323e-02 -7.457241e-03
#> Locus_2739 2.321087e-03 -2.325296e-02 -2.502939e-02 8.993651e-02
#> Locus_2753 -1.093686e-02 -2.212321e-02 1.113734e-02 1.281375e-01
#> Locus_2755 2.046984e-02 3.578494e-03 -6.217066e-03 1.759591e-02
#> Locus_2758 2.045566e-01 -1.692100e-01 6.761348e-02 -1.230663e-01
#> Locus_2760 3.506933e-03 8.741992e-02 -1.939889e-02 4.540998e-03
#> Locus_2761 -6.262224e-03 6.276579e-02 -2.204298e-02 1.599476e-02
#> Locus_2764 -5.325116e-02 -2.944157e-02 -1.442323e-02 -7.457241e-03
#> Locus_2766 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2767 9.016616e-02 -1.812956e-01 -1.210801e-01 -7.995000e-02
#> Locus_2768 -1.550082e-02 3.153482e-02 -2.057255e-02 -8.011266e-03
#> Locus_2770 -6.132617e-02 -4.775015e-02 7.277528e-03 -4.039450e-02
#> Locus_2774 -4.317496e-02 -3.618608e-02 -6.204467e-03 -3.379137e-02
#> Locus_2776 1.858304e-02 1.103878e-02 -1.316149e-02 1.995691e-03
#> Locus_2789 1.750415e-02 -3.029431e-02 8.536520e-02 4.310297e-01
#> Locus_2792 1.560602e-03 1.698532e-02 -1.273945e-02 -8.658562e-03
#> Locus_2793 3.222528e-02 -1.504975e-02 2.616025e-02 -2.390265e-02
#> Locus_2794 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2795 -1.154108e-01 1.441891e-03 2.250240e-02 2.500903e-01
#> Locus_2796 -2.542117e-02 -1.520932e-02 5.559921e-03 2.628821e-02
#> Locus_2798 -1.204379e-02 2.102857e-02 3.709140e-02 -1.975608e-02
#> Locus_2804 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2806 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2808 2.517740e-02 -6.687043e-03 1.486889e-03 -9.963326e-03
#> Locus_2809 -1.986935e-02 -1.482855e-02 5.189881e-03 -1.174941e-02
#> Locus_2810 3.677561e-02 2.509829e-02 -3.557386e-02 -3.168927e-02
#> Locus_2811 -2.564650e-02 2.078181e-03 2.017368e-02 -2.537829e-02
#> Locus_2814 -6.420917e-02 -2.965011e-02 5.409830e-03 1.930089e-02
#> Locus_2816 -2.158748e-02 -1.809304e-02 -3.102234e-03 -1.689569e-02
#> Locus_2817 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2818 -1.328294e-01 -5.456637e-02 -5.357523e-03 -3.919049e-02
#> Locus_2823 2.386589e-01 -1.591190e-02 2.114367e-02 -1.678459e-01
#> Locus_2830 -2.942508e-02 2.867446e-02 -1.473878e-02 -3.319890e-02
#> Locus_2831 2.738938e-01 1.763708e-02 9.198610e-02 4.444428e-02
#> Locus_2832 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2835 2.895363e-02 -1.216440e-02 -2.430351e-02 -1.723377e-02
#> Locus_2836 -4.156818e-02 -1.930449e-02 -3.636007e-03 -6.986679e-03
#> Locus_2839 1.361826e-01 -5.593238e-02 -2.188911e-01 -1.380698e-01
#> Locus_2842 -6.655728e-02 1.303402e-01 -1.493594e-02 4.079593e-01
#> Locus_2844 -2.114233e-02 -8.168065e-03 3.132299e-02 8.577284e-02
#> Locus_2845 3.102574e-03 4.963783e-02 3.221601e-02 -2.791766e-02
#> Locus_2850 -5.960804e-02 -4.448566e-02 1.556964e-02 -3.078063e-02
#> Locus_2852 3.090640e-02 1.641438e-02 2.718429e-02 1.486121e-01
#> Locus_2854 -1.494260e-02 -4.583701e-03 3.575609e-03 -7.261874e-03
#> Locus_2856 -1.705937e-02 4.774584e-02 2.860109e-02 -9.320936e-02
#> Locus_2861 -1.195924e-02 1.019087e-01 5.993032e-02 1.664629e-01
#> Locus_2865 -2.881668e-02 -1.361011e-02 -9.766868e-03 -6.732373e-03
#> Locus_2868 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2871 2.321087e-03 -2.325296e-02 -2.502939e-02 8.993651e-02
#> Locus_2872 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2873 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2874 -1.254989e-02 7.354897e-02 -5.003577e-03 8.603369e-02
#> Locus_2875 -8.333460e-04 -1.670290e-02 -3.339391e-03 6.629651e-02
#> Locus_2878 -1.504805e-03 6.581785e-03 1.019391e-02 6.687764e-02
#> Locus_2881 -8.873857e-03 -2.051960e-05 8.015739e-04 -1.102771e-02
#> Locus_2884 -4.853913e-02 -2.711542e-02 7.249782e-03 9.315332e-02
#> Locus_2890 -1.960520e-03 -1.328179e-02 -5.558796e-03 6.282795e-02
#> Locus_2895 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2896 5.991734e-02 -5.835502e-02 2.563608e-02 -4.256361e-02
#> Locus_2904 -1.564025e-01 -5.587293e-02 3.221545e-02 -3.475885e-02
#> Locus_2905 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2910 -9.069416e-03 1.248427e-02 3.706344e-02 -1.290967e-02
#> Locus_2915 1.081166e-01 -1.400226e-01 -2.055909e-05 -1.000067e-01
#> Locus_2916 1.278587e-02 -7.031674e-02 -5.769852e-02 -2.253500e-02
#> Locus_2923 -1.929842e-02 6.521978e-03 -1.608385e-02 -1.753075e-02
#> Locus_2932 -4.195692e-02 7.553012e-02 5.183928e-02 6.223117e-02
#> Locus_2933 5.477085e-03 -1.031841e-02 8.425221e-03 6.086581e-02
#> Locus_2934 -8.189157e-02 -2.964553e-02 9.355890e-03 -2.193098e-02
#> Locus_2936 1.012554e-02 -2.490716e-02 4.780413e-02 -5.785592e-02
#> Locus_2937 -2.674479e-03 3.413536e-02 7.658193e-02 4.513230e-01
#> Locus_2938 -7.905428e-02 -4.203371e-02 -1.127601e-03 -2.642572e-02
#> Locus_2947 -3.589864e-01 -1.886685e-01 -3.255616e-02 -8.822165e-02
#> Locus_2949 1.832347e-02 -2.074730e-02 5.158049e-03 9.194628e-02
#> Locus_2951 -1.468925e-02 -1.254714e-03 -9.090753e-05 1.591877e-01
#> Locus_2953 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2954 -5.155980e-04 -1.768886e-03 3.730178e-03 -1.978950e-02
#> Locus_2962 6.139458e-02 -8.081219e-02 -2.788994e-02 -3.040403e-02
#> Locus_2966 7.695414e-03 5.951227e-02 3.411161e-03 -5.269969e-02
#> Locus_2972 1.351818e-01 -1.496057e-01 -2.156295e-02 -7.753374e-02
#> Locus_2974 4.605800e-03 5.977836e-02 9.970063e-03 8.674240e-02
#> Locus_2976 7.480669e-03 3.011761e-02 -2.182228e-02 -2.235630e-02
#> Locus_2977 9.350097e-02 -6.077448e-02 4.497874e-02 -4.300976e-02
#> Locus_2978 -3.458431e-02 7.592757e-03 5.465418e-02 6.349266e-02
#> Locus_2983 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2991 -9.432081e-03 3.609800e-02 -2.334658e-02 -1.177710e-02
#> Locus_2993 -1.467035e-01 -6.359278e-02 -1.870000e-02 -3.866099e-02
#> Locus_2994 -1.586816e-02 -9.170355e-03 2.649791e-02 -2.301579e-02
#> Locus_2995 -1.252445e-02 1.255316e-01 -4.408595e-02 3.198953e-02
#> Locus_3005 2.323293e-02 -1.026720e-04 4.285258e-02 -1.890992e-02
#> Locus_3009 -3.585417e-02 -9.524296e-03 -1.279099e-02 -1.157573e-02
#> Locus_3010 6.921294e-02 3.777474e-02 -7.719988e-02 2.900436e-01
#> Locus_3012 6.957269e-02 1.080710e-01 -1.932214e-02 4.574516e-01
#> Locus_3013 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3023 4.117779e-02 -1.719492e-02 3.549517e-02 -4.241752e-02
#> Locus_3029 6.342161e-04 9.099577e-03 2.278844e-02 1.339174e-01
#> Locus_3030 4.282312e-02 -8.628943e-02 -4.398678e-02 -3.809340e-02
#> Locus_3032 -8.848468e-02 -4.524291e-02 -1.978618e-02 -1.320049e-02
#> Locus_3042 -4.102963e-03 1.666175e-02 3.195588e-02 6.024196e-02
#> Locus_3043 9.052891e-03 7.616535e-02 2.666424e-02 7.892649e-02
#> Locus_3046 -4.657210e-03 4.242478e-02 2.418596e-03 9.665920e-03
#> Locus_3047 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3051 -9.378967e-03 -4.611419e-03 7.448867e-02 1.834452e-01
#> Locus_3054 -1.553615e-02 1.441107e-02 4.671075e-02 1.305175e-02
#> Locus_3056 2.843853e-02 -6.395585e-02 -3.180762e-02 -3.710911e-02
#> Locus_3058 1.622008e-01 -1.617329e-01 -5.553932e-03 -2.466186e-01
#> Locus_3059 4.903414e-02 3.346439e-02 -4.743181e-02 -4.220392e-02
#> Locus_3060 3.116073e-02 -2.367179e-02 4.356049e-03 -1.194097e-02
#> Locus_3061 4.384021e-02 -1.944311e-03 -9.744804e-03 -1.866679e-02
#> Locus_3062 -3.049306e-02 -1.952014e-02 1.448716e-02 -5.534520e-03
#> Locus_3063 -2.123247e-02 1.201256e-01 4.897246e-02 2.098986e-01
#> Locus_3064 -9.393336e-03 9.414869e-02 -3.306446e-02 2.447845e-02
#> Locus_3069 1.159418e-01 3.934246e-02 -3.226686e-02 -3.578459e-03
#> Locus_3078 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3079 -4.102963e-03 1.666175e-02 3.195588e-02 6.024196e-02
#> Locus_3081 -8.333460e-04 -1.670290e-02 -3.339391e-03 6.629651e-02
#> Locus_3084 1.118284e-02 4.675648e-02 1.183917e-02 9.920704e-02
#> Locus_3087 -2.063968e-02 4.529402e-02 -4.228916e-02 -7.785641e-02
#> Locus_3089 -5.750088e-02 3.093221e-02 8.033002e-05 -4.069398e-02
#> Locus_3090 -8.555719e-03 3.509288e-03 2.002877e-02 1.709782e-02
#> Locus_3091 3.252499e-02 -9.547251e-02 -3.680077e-02 6.627987e-02
#> Locus_3092 -4.752946e-02 -4.289737e-02 -9.279811e-03 -2.102393e-02
#> Locus_3093 7.377955e-03 -2.149050e-02 2.301496e-03 9.440708e-02
#> Locus_3097 2.134327e-02 7.225468e-03 -3.332978e-02 -4.327155e-03
#> Locus_3098 -9.902985e-03 -2.754887e-02 3.305956e-02 1.523350e-01
#> Locus_3102 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3116 -4.145683e-02 -3.292160e-02 2.087647e-03 -2.864509e-02
#> Locus_3122 2.271837e-01 -1.902764e-02 2.154310e-02 -3.504046e-01
#> Locus_3126 5.772637e-02 -1.254927e-02 8.607027e-02 -4.278662e-02
#> Locus_3128 8.964627e-03 1.084206e-01 4.265831e-02 5.240486e-01
#> Locus_3129 2.287699e-02 -1.239526e-02 2.179140e-02 -1.622310e-02
#> Locus_3133 1.813883e-02 -2.496854e-02 -7.412689e-02 2.581935e-02
#> Locus_3138 7.816070e-03 8.018554e-02 1.579409e-02 -2.882655e-02
#> Locus_3140 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3143 -1.277361e-01 -6.523808e-02 -5.183928e-02 -7.662033e-02
#> Locus_3145 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3147 8.527949e-02 -5.068741e-04 -2.641646e-03 -6.203914e-02
#> Locus_3148 -8.244218e-02 1.282587e-03 -6.209953e-03 -1.208143e-02
#> Locus_3154 2.049871e-02 -5.633474e-03 4.563692e-02 1.900766e-02
#> Locus_3155 1.161646e-02 -5.133599e-05 2.142629e-02 -1.166507e-02
#> Locus_3160 -1.973852e-01 -8.404777e-02 7.959524e-03 -3.938580e-02
#> Locus_3161 7.891866e-02 -4.392819e-03 6.303251e-02 -3.984086e-02
#> Locus_3164 -1.165864e-04 9.628077e-02 -4.503069e-02 2.095056e-01
#> Locus_3165 1.136047e-02 7.938393e-03 -3.157820e-02 2.124195e-02
#> Locus_3167 3.166253e-02 -8.570031e-03 -1.808418e-03 -1.776912e-02
#> Locus_3168 -7.699518e-03 3.646732e-02 1.849369e-02 2.404153e-02
#> Locus_3169 -1.107083e-01 -4.325565e-02 -4.109783e-04 -3.348137e-02
#> Locus_3171 2.983582e-02 7.151873e-03 4.267885e-02 -1.740332e-02
#> Locus_3175 1.993917e-02 -3.317540e-02 1.414820e-02 1.760077e-01
#> Locus_3177 6.010060e-02 7.988955e-02 -2.483393e-02 -1.885370e-01
#> Locus_3179 -1.265173e-01 -4.670553e-02 2.506423e-02 -2.987112e-02
#> Locus_3181 2.317242e-02 -2.605265e-02 -1.175134e-02 2.762242e-02
#> Locus_3182 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3188 1.227668e-02 8.579829e-02 5.883581e-02 2.060651e-01
#> Locus_3190 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3191 2.279810e-03 -2.013521e-02 -1.345391e-02 -1.783852e-02
#> Locus_3192 -1.314227e-02 1.078952e-01 -3.116758e-02 -4.538444e-02
#> Locus_3197 -1.019787e-01 9.451924e-02 -3.097190e-02 2.987667e-03
#> Locus_3198 -4.424234e-02 -2.262146e-02 -9.893090e-03 -1.782238e-03
#> PC2 PC3
#> Locus_10 1.511513e-02 3.391936e-02
#> Locus_15 9.031505e-03 1.795886e-02
#> Locus_22 -2.410855e-02 -8.720620e-03
#> Locus_28 3.464025e-02 4.337825e-02
#> Locus_32 7.482888e-03 2.772942e-02
#> Locus_35 -1.174114e-16 4.451135e-17
#> Locus_37 -6.455618e-17 -1.543825e-17
#> Locus_61 -2.588908e-01 -2.243204e-01
#> Locus_71 -3.021704e-02 -4.008441e-03
#> Locus_72 3.053708e-17 -2.172114e-17
#> Locus_82 -5.775486e-18 -4.080647e-17
#> Locus_84 -1.655639e-02 1.238370e-02
#> Locus_90 4.839862e-02 1.564788e-01
#> Locus_91 5.542628e-02 1.236429e-01
#> Locus_96 -4.923400e-17 -3.689745e-17
#> Locus_104 -2.098904e-02 1.859855e-03
#> Locus_122 -3.586408e-02 -6.545240e-02
#> Locus_125 3.190852e-02 -3.371845e-02
#> Locus_131 -7.017408e-02 -7.009538e-03
#> Locus_133 5.551278e-17 -2.289696e-17
#> Locus_138 -8.631762e-02 -1.427025e-01
#> Locus_154 3.437187e-02 -7.270074e-02
#> Locus_162 9.912627e-02 -2.313426e-02
#> Locus_166 8.981252e-02 1.805598e-01
#> Locus_177 -8.073273e-18 1.380687e-17
#> Locus_185 9.151479e-17 -8.686149e-17
#> Locus_191 5.577431e-02 4.683311e-02
#> Locus_193 -2.589389e-01 -7.156164e-02
#> Locus_207 -2.231504e-02 -1.162235e-01
#> Locus_211 1.786337e-02 4.810218e-02
#> Locus_213 -1.431080e-01 -1.491599e-02
#> Locus_214 3.997398e-02 1.065591e-01
#> Locus_222 -5.972515e-03 2.153054e-02
#> Locus_227 -1.868714e-01 -1.010299e-01
#> Locus_242 2.884387e-02 2.613665e-02
#> Locus_249 -1.078396e-01 -1.935897e-01
#> Locus_254 1.219059e-02 -1.685985e-02
#> Locus_256 -5.190816e-02 -1.675669e-01
#> Locus_263 -5.249508e-02 2.506512e-01
#> Locus_268 -1.326819e-02 -3.230225e-03
#> Locus_279 7.533742e-03 5.831542e-02
#> Locus_285 -1.302212e-01 -4.602961e-02
#> Locus_287 6.851495e-02 -5.134228e-02
#> Locus_292 8.198562e-03 -4.019037e-03
#> Locus_294 3.273241e-02 -6.377052e-04
#> Locus_303 1.938389e-17 3.882605e-17
#> Locus_308 -2.077013e-02 1.396465e-02
#> Locus_312 5.410733e-02 2.154704e-02
#> Locus_313 8.207201e-20 1.137034e-17
#> Locus_314 4.964633e-02 4.797686e-02
#> Locus_317 1.098179e-03 -1.034085e-02
#> Locus_318 5.205393e-02 -1.768178e-01
#> Locus_323 0.000000e+00 0.000000e+00
#> Locus_330 -1.401990e-02 -4.087007e-03
#> Locus_331 7.108136e-04 -1.127733e-02
#> Locus_333 1.361199e-02 3.152316e-02
#> Locus_334 2.244301e-02 1.527092e-02
#> Locus_336 2.929516e-02 -3.198850e-02
#> Locus_337 2.172907e-02 3.597516e-02
#> Locus_339 -6.112741e-02 -3.092584e-03
#> Locus_341 3.395697e-02 4.997430e-02
#> Locus_345 -2.796431e-02 -6.129038e-03
#> Locus_350 -4.563590e-02 -1.162436e-02
#> Locus_354 6.854008e-04 -2.201722e-02
#> Locus_359 -3.640473e-01 -8.380259e-02
#> Locus_365 -1.081279e-02 -1.347147e-02
#> Locus_366 -1.500952e-02 -1.226637e-02
#> Locus_369 2.390637e-02 -2.675838e-02
#> Locus_372 -4.068941e-03 5.263743e-04
#> Locus_389 -4.253641e-02 2.641598e-02
#> Locus_391 1.963806e-02 5.226529e-02
#> Locus_392 5.015911e-02 1.544315e-01
#> Locus_393 -2.367477e-02 1.116977e-02
#> Locus_394 0.000000e+00 0.000000e+00
#> Locus_396 -1.566540e-02 -9.709681e-03
#> Locus_399 4.808343e-02 1.063108e-01
#> Locus_400 9.013908e-03 1.522143e-02
#> Locus_401 1.170684e-01 3.869810e-02
#> Locus_404 -9.192274e-02 -5.767279e-02
#> Locus_406 2.310110e-01 -1.534971e-01
#> Locus_409 -2.303192e-05 -1.045968e-02
#> Locus_420 -1.098763e-02 -1.817652e-02
#> Locus_422 -5.239500e-02 2.308996e-02
#> Locus_427 -6.156339e-02 -6.917301e-02
#> Locus_430 0.000000e+00 0.000000e+00
#> Locus_436 -3.616699e-02 3.893238e-02
#> Locus_438 -2.448848e-02 -8.593796e-03
#> Locus_439 9.576799e-03 -9.036716e-03
#> Locus_441 -2.300517e-01 -1.202631e-01
#> Locus_445 -2.598617e-02 -3.430636e-04
#> Locus_448 -1.755876e-02 -6.059028e-03
#> Locus_449 -3.685496e-02 -2.646124e-02
#> Locus_456 -2.245618e-02 1.036163e-02
#> Locus_457 -2.050055e-02 1.953570e-02
#> Locus_458 -2.701149e-02 -6.772018e-02
#> Locus_460 0.000000e+00 0.000000e+00
#> Locus_461 -5.948436e-02 6.313117e-03
#> Locus_462 0.000000e+00 0.000000e+00
#> Locus_464 1.822047e-02 -3.211892e-03
#> Locus_469 -3.059126e-02 1.876182e-01
#> Locus_475 5.279073e-02 1.092828e-02
#> Locus_480 7.054833e-03 1.740156e-01
#> Locus_482 2.703482e-02 -2.387164e-02
#> Locus_483 1.426711e-02 6.664209e-02
#> Locus_491 2.304563e-02 -1.931420e-02
#> Locus_498 -6.645343e-02 3.305217e-03
#> Locus_500 -7.564493e-03 -1.653190e-03
#> Locus_503 -9.381254e-02 -4.417257e-02
#> Locus_505 -5.298189e-02 -3.865849e-04
#> Locus_508 -1.273356e-01 -1.746192e-01
#> Locus_512 2.487013e-02 -2.728533e-02
#> Locus_516 2.650123e-03 1.657940e-02
#> Locus_517 9.743810e-02 1.497557e-02
#> Locus_518 -5.769606e-02 -1.376867e-02
#> Locus_528 0.000000e+00 0.000000e+00
#> Locus_529 -3.456662e-02 -7.355444e-02
#> Locus_531 1.743021e-02 1.139010e-01
#> Locus_532 -4.809406e-03 3.120380e-02
#> Locus_533 3.489741e-02 -2.909943e-02
#> Locus_535 -1.498448e-02 1.293378e-02
#> Locus_546 -6.121942e-02 1.264082e-02
#> Locus_557 0.000000e+00 0.000000e+00
#> Locus_558 -5.537367e-02 -9.143174e-02
#> Locus_561 0.000000e+00 0.000000e+00
#> Locus_562 -1.109110e-01 2.412713e-03
#> Locus_564 -3.521417e-02 -6.211359e-03
#> Locus_566 1.671037e-02 1.752034e-02
#> Locus_572 -1.412081e-01 -3.502985e-02
#> Locus_573 -6.017982e-02 -2.603701e-03
#> Locus_579 -6.762156e-02 2.772551e-02
#> Locus_581 -2.186871e-01 -5.606089e-02
#> Locus_582 -6.017982e-02 -2.603701e-03
#> Locus_583 4.755154e-02 8.316848e-02
#> Locus_586 -1.320673e-01 -1.776315e-02
#> Locus_589 -2.445006e-01 -1.309120e-01
#> Locus_596 -3.398277e-02 -9.240698e-03
#> Locus_597 8.122935e-03 -3.201450e-02
#> Locus_598 0.000000e+00 0.000000e+00
#> Locus_600 -1.497894e-02 -1.298688e-02
#> Locus_602 -6.036119e-02 7.841606e-03
#> Locus_603 0.000000e+00 0.000000e+00
#> Locus_604 9.454631e-03 -1.829443e-03
#> Locus_609 1.729876e-02 7.698372e-02
#> Locus_612 5.644543e-02 1.292286e-02
#> Locus_613 5.426301e-03 4.576095e-02
#> Locus_614 -3.252449e-02 -9.009799e-03
#> Locus_622 1.109244e-02 -5.935010e-03
#> Locus_625 2.369553e-02 -7.702403e-03
#> Locus_626 4.609125e-02 -3.862840e-02
#> Locus_633 -8.461991e-03 -3.470303e-03
#> Locus_634 -1.728331e-02 -3.677722e-02
#> Locus_635 -2.096097e-01 9.735391e-02
#> Locus_636 -9.879419e-02 2.446689e-01
#> Locus_637 -3.670486e-02 -6.536052e-02
#> Locus_639 -2.893731e-02 -9.991127e-03
#> Locus_642 0.000000e+00 0.000000e+00
#> Locus_647 2.884813e-02 -6.082043e-02
#> Locus_652 -5.324072e-03 -2.824365e-02
#> Locus_665 8.896506e-03 1.461888e-02
#> Locus_671 9.114145e-02 -7.124409e-02
#> Locus_679 -2.052159e-02 -2.463376e-02
#> Locus_681 -9.994584e-03 -3.713030e-02
#> Locus_682 9.114145e-02 -7.124409e-02
#> Locus_683 0.000000e+00 0.000000e+00
#> Locus_685 -6.017982e-02 -2.603701e-03
#> Locus_687 8.838931e-03 4.563831e-03
#> Locus_691 1.877589e-03 -1.466048e-03
#> Locus_699 1.884072e-02 1.541533e-01
#> Locus_701 0.000000e+00 0.000000e+00
#> Locus_707 1.254816e-02 8.827872e-02
#> Locus_710 2.612277e-03 6.513358e-02
#> Locus_711 1.133735e-02 6.517075e-03
#> Locus_712 0.000000e+00 0.000000e+00
#> Locus_717 -2.052159e-02 -2.463376e-02
#> Locus_718 0.000000e+00 0.000000e+00
#> Locus_724 1.794686e-02 8.550223e-03
#> Locus_725 -1.728331e-02 -3.677722e-02
#> Locus_727 1.956770e-02 3.340087e-02
#> Locus_729 0.000000e+00 0.000000e+00
#> Locus_733 1.065048e-01 -9.727007e-02
#> Locus_734 -1.906536e-02 -3.336578e-03
#> Locus_735 -6.806754e-02 -6.774555e-02
#> Locus_736 9.114145e-02 -7.124409e-02
#> Locus_740 3.195062e-01 1.808189e-02
#> Locus_742 -6.344401e-03 8.226648e-03
#> Locus_746 -2.277150e-02 5.403997e-02
#> Locus_750 -2.974218e-02 3.156558e-03
#> Locus_751 2.218489e-02 -1.187002e-02
#> Locus_753 -1.800878e-02 2.686218e-02
#> Locus_758 -1.942154e-02 -2.858330e-02
#> Locus_765 -1.219710e-02 -2.325883e-02
#> Locus_769 -2.581582e-02 -2.396298e-02
#> Locus_770 -5.069832e-02 3.383281e-01
#> Locus_771 1.415655e-01 -1.616278e-01
#> Locus_776 0.000000e+00 0.000000e+00
#> Locus_779 2.570294e-02 2.213075e-02
#> Locus_784 2.382504e-02 -1.043940e-02
#> Locus_786 9.519999e-02 -8.435246e-02
#> Locus_787 3.371682e-02 -5.322080e-04
#> Locus_798 -1.268064e-01 4.660009e-03
#> Locus_802 7.500742e-02 -7.650215e-02
#> Locus_803 -7.746313e-02 -3.938092e-02
#> Locus_807 -5.948436e-02 6.313117e-03
#> Locus_808 -8.997989e-03 -2.243963e-02
#> Locus_809 2.218489e-02 -1.187002e-02
#> Locus_811 -4.034889e-02 -5.111902e-03
#> Locus_816 2.080870e-02 3.553624e-03
#> Locus_821 -6.455412e-03 -2.433817e-03
#> Locus_825 -5.140581e-02 -7.080719e-02
#> Locus_831 -1.616423e-01 7.429126e-03
#> Locus_835 1.182243e-01 -7.008675e-02
#> Locus_838 1.043319e-01 -1.006075e-01
#> Locus_847 4.328394e-02 -1.672914e-02
#> Locus_850 3.219924e-02 1.322384e-02
#> Locus_851 3.030926e-02 7.104728e-02
#> Locus_853 0.000000e+00 0.000000e+00
#> Locus_854 -2.649095e-02 -1.932925e-04
#> Locus_859 3.190852e-02 -3.371845e-02
#> Locus_861 2.790774e-02 2.419749e-02
#> Locus_864 7.211461e-02 1.918499e-01
#> Locus_869 8.121681e-02 -8.387880e-02
#> Locus_870 -1.501018e-01 -2.050843e-03
#> Locus_875 0.000000e+00 0.000000e+00
#> Locus_876 -9.063860e-02 -1.640313e-03
#> Locus_879 -1.527833e-01 -2.685294e-02
#> Locus_881 -1.336156e-01 1.871926e-01
#> Locus_887 2.385919e-02 3.193290e-02
#> Locus_888 0.000000e+00 0.000000e+00
#> Locus_895 1.109519e-02 3.715425e-02
#> Locus_896 6.653198e-02 2.025658e-01
#> Locus_897 6.300292e-03 -4.396091e-03
#> Locus_898 -9.774135e-02 -4.879840e-02
#> Locus_899 5.831762e-02 -7.453220e-02
#> Locus_901 -1.004065e-01 -1.263396e-01
#> Locus_902 1.574223e-01 -1.649247e-01
#> Locus_903 -1.765928e-01 -2.244135e-03
#> Locus_905 -5.395074e-02 1.894539e-02
#> Locus_908 0.000000e+00 0.000000e+00
#> Locus_909 -7.576968e-03 -2.043111e-02
#> Locus_915 0.000000e+00 0.000000e+00
#> Locus_924 -2.661559e-02 3.825824e-02
#> Locus_932 -2.949727e-02 1.560864e-02
#> Locus_935 0.000000e+00 0.000000e+00
#> Locus_936 -4.329324e-02 1.261467e-02
#> Locus_938 -3.456662e-02 -7.355444e-02
#> Locus_941 0.000000e+00 0.000000e+00
#> Locus_944 -4.744556e-03 5.000632e-02
#> Locus_945 2.154719e-01 -1.906567e-01
#> Locus_946 0.000000e+00 0.000000e+00
#> Locus_949 8.914643e-03 -4.170400e-03
#> Locus_950 -1.299959e-01 2.501077e-02
#> Locus_951 5.851632e-02 2.494921e-01
#> Locus_953 -3.498002e-02 -1.742469e-03
#> Locus_958 0.000000e+00 0.000000e+00
#> Locus_965 -3.581240e-02 2.568591e-02
#> Locus_966 3.742165e-02 8.451591e-02
#> Locus_967 9.390857e-03 5.185556e-02
#> Locus_968 -3.495209e-02 4.542185e-02
#> Locus_970 -7.256297e-02 -3.762012e-02
#> Locus_972 -1.234515e-02 -1.777255e-02
#> Locus_974 -8.992200e-02 5.528576e-04
#> Locus_979 -1.741615e-02 -6.626832e-04
#> Locus_992 -6.547529e-03 -8.796063e-03
#> Locus_998 4.739105e-02 -1.540481e-02
#> Locus_999 1.926437e-03 1.247592e-02
#> Locus_1002 -7.549692e-03 -2.564351e-02
#> Locus_1005 7.969465e-02 2.622142e-01
#> Locus_1006 -6.742174e-03 1.797006e-03
#> Locus_1007 7.605701e-02 8.916155e-02
#> Locus_1015 2.350939e-02 9.118807e-02
#> Locus_1016 0.000000e+00 0.000000e+00
#> Locus_1017 2.413466e-02 -6.259413e-02
#> Locus_1023 -1.728331e-02 -3.677722e-02
#> Locus_1024 2.075684e-02 1.068474e-02
#> Locus_1025 -6.522002e-03 2.329413e-02
#> Locus_1027 7.038391e-03 -6.433120e-03
#> Locus_1030 -3.551095e-02 2.436720e-01
#> Locus_1031 1.215175e-02 7.483146e-02
#> Locus_1033 -2.378033e-02 -3.611979e-02
#> Locus_1034 4.597803e-02 -8.535201e-03
#> Locus_1035 1.778359e-02 3.225062e-02
#> Locus_1038 9.454631e-03 -1.829443e-03
#> Locus_1041 -4.221482e-02 2.124008e-02
#> Locus_1042 3.012644e-03 3.221361e-02
#> Locus_1044 -6.667868e-02 -8.553232e-03
#> Locus_1046 8.915869e-03 -1.042610e-02
#> Locus_1051 1.595426e-02 -1.685922e-02
#> Locus_1054 -1.727330e-02 -1.281548e-02
#> Locus_1055 0.000000e+00 0.000000e+00
#> Locus_1065 1.009517e-02 9.286300e-02
#> Locus_1071 -3.884309e-02 -5.716661e-02
#> Locus_1072 -1.675579e-02 -3.642744e-03
#> Locus_1073 1.400977e-02 -3.083353e-02
#> Locus_1074 2.941747e-02 4.712015e-02
#> Locus_1075 -4.993758e-02 2.101836e-02
#> Locus_1076 0.000000e+00 0.000000e+00
#> Locus_1080 -1.657250e-02 -4.805455e-02
#> Locus_1082 0.000000e+00 0.000000e+00
#> Locus_1093 -3.620093e-02 -7.679444e-03
#> Locus_1095 0.000000e+00 0.000000e+00
#> Locus_1098 -2.581582e-02 -2.396298e-02
#> Locus_1099 2.929516e-02 -3.198850e-02
#> Locus_1105 -5.800806e-02 -9.344989e-02
#> Locus_1106 -1.589962e-02 2.666565e-02
#> Locus_1107 -1.345912e-02 -5.673221e-03
#> Locus_1108 -1.203596e-01 -5.207401e-03
#> Locus_1109 0.000000e+00 0.000000e+00
#> Locus_1113 -5.034315e-02 -9.517740e-03
#> Locus_1115 0.000000e+00 0.000000e+00
#> Locus_1118 4.680804e-02 -5.160631e-02
#> Locus_1122 0.000000e+00 0.000000e+00
#> Locus_1127 -6.017982e-02 -2.603701e-03
#> Locus_1138 0.000000e+00 0.000000e+00
#> Locus_1141 -1.671645e-01 -2.731053e-02
#> Locus_1143 2.209726e-02 1.406141e-01
#> Locus_1144 -3.299180e-02 -8.894267e-03
#> Locus_1155 -7.497833e-03 -3.277463e-02
#> Locus_1158 7.014541e-02 2.521834e-02
#> Locus_1159 0.000000e+00 0.000000e+00
#> Locus_1161 2.675988e-02 6.272105e-02
#> Locus_1162 1.426212e-02 1.162149e-01
#> Locus_1166 1.874628e-02 2.833816e-02
#> Locus_1172 -1.417742e-01 -4.222469e-02
#> Locus_1179 0.000000e+00 0.000000e+00
#> Locus_1181 8.915869e-03 -1.042610e-02
#> Locus_1186 -6.693919e-02 -1.074153e-01
#> Locus_1191 1.192371e-02 -4.191330e-03
#> Locus_1197 -4.734954e-02 2.233953e-02
#> Locus_1199 -2.908570e-02 -2.981827e-02
#> Locus_1201 -8.276351e-02 -4.897912e-02
#> Locus_1204 -1.326819e-02 -3.230225e-03
#> Locus_1206 2.382504e-02 -1.043940e-02
#> Locus_1211 -1.108253e-01 8.273247e-03
#> Locus_1212 -1.127708e-02 -1.599367e-02
#> Locus_1217 -6.002784e-02 -1.330048e-02
#> Locus_1223 -1.755876e-02 -6.059028e-03
#> Locus_1224 1.261405e-02 1.778835e-02
#> Locus_1227 1.670517e-02 -2.306628e-02
#> Locus_1231 -1.203596e-01 -5.207401e-03
#> Locus_1235 0.000000e+00 0.000000e+00
#> Locus_1236 -7.569411e-02 -7.736913e-02
#> Locus_1239 1.023657e-01 -1.300904e-01
#> Locus_1247 6.178684e-02 -9.375213e-02
#> Locus_1248 0.000000e+00 0.000000e+00
#> Locus_1249 0.000000e+00 0.000000e+00
#> Locus_1250 -3.557342e-02 6.093777e-03
#> Locus_1254 -8.992200e-02 5.528576e-04
#> Locus_1255 3.341469e-03 -1.013177e-02
#> Locus_1256 -4.891490e-02 -4.941533e-02
#> Locus_1258 -2.235115e-01 -1.327719e-01
#> Locus_1262 2.074164e-02 1.073113e-01
#> Locus_1263 1.103604e-01 1.965499e-02
#> Locus_1265 -1.559066e-02 -3.263084e-03
#> Locus_1266 -1.539850e-02 -6.128303e-03
#> Locus_1270 -2.009387e-02 -4.179970e-03
#> Locus_1278 2.380339e-01 -2.064377e-01
#> Locus_1279 8.558623e-02 -6.292831e-02
#> Locus_1284 8.236869e-02 -5.678153e-02
#> Locus_1286 -1.355259e-01 -5.562247e-02
#> Locus_1287 -3.687027e-01 -1.756822e-01
#> Locus_1290 -1.728331e-02 -3.677722e-02
#> Locus_1295 5.656702e-04 -1.290492e-02
#> Locus_1301 4.714326e-03 8.432274e-02
#> Locus_1302 -4.791078e-02 -3.312303e-02
#> Locus_1305 4.152529e-02 7.450024e-04
#> Locus_1313 -3.946434e-03 1.328646e-02
#> Locus_1315 0.000000e+00 0.000000e+00
#> Locus_1316 -5.950497e-02 -3.132015e-02
#> Locus_1318 2.673290e-01 -2.384262e-01
#> Locus_1319 -1.798440e-01 1.105715e-03
#> Locus_1338 7.806299e-02 5.533232e-02
#> Locus_1343 -1.297694e-01 4.555639e-02
#> Locus_1345 -2.056743e-02 -1.288307e-02
#> Locus_1347 -3.013018e-02 8.735949e-03
#> Locus_1353 -8.943565e-03 1.108354e-02
#> Locus_1354 2.473462e-02 4.434918e-02
#> Locus_1355 4.121887e-02 -3.617983e-02
#> Locus_1360 0.000000e+00 0.000000e+00
#> Locus_1363 0.000000e+00 0.000000e+00
#> Locus_1364 -1.511764e-02 7.523211e-03
#> Locus_1366 1.829653e-02 1.551578e-02
#> Locus_1370 -1.769625e-01 -1.820323e-01
#> Locus_1374 1.807384e-01 -1.962726e-01
#> Locus_1376 -1.113928e-01 2.827590e-02
#> Locus_1377 -2.084851e-02 -2.982245e-02
#> Locus_1379 3.724458e-03 6.164508e-02
#> Locus_1380 0.000000e+00 0.000000e+00
#> Locus_1382 -3.895089e-02 -3.960215e-02
#> Locus_1384 4.538536e-02 -2.842091e-02
#> Locus_1392 1.337215e-02 1.413346e-01
#> Locus_1396 -1.147952e-01 -3.568946e-02
#> Locus_1399 0.000000e+00 0.000000e+00
#> Locus_1404 0.000000e+00 0.000000e+00
#> Locus_1408 -7.067857e-02 -8.018215e-03
#> Locus_1414 -6.903431e-02 -1.059253e-01
#> Locus_1416 -2.263976e-03 5.034704e-02
#> Locus_1420 -2.392137e-02 3.675956e-02
#> Locus_1423 -7.459515e-02 -7.435823e-02
#> Locus_1424 -2.041097e-02 3.067669e-02
#> Locus_1425 -1.611860e-02 -1.148574e-02
#> Locus_1426 7.658075e-02 1.912887e-01
#> Locus_1427 1.109244e-02 -5.935010e-03
#> Locus_1430 -1.908099e-02 -8.787670e-02
#> Locus_1433 -1.080911e-01 -3.398949e-02
#> Locus_1439 1.061839e-01 -5.141523e-02
#> Locus_1447 -5.081052e-03 2.520602e-02
#> Locus_1452 7.930690e-02 -6.678561e-02
#> Locus_1455 -1.750413e-01 1.667928e-01
#> Locus_1457 0.000000e+00 0.000000e+00
#> Locus_1459 -1.021579e-01 1.116010e-03
#> Locus_1460 -4.328526e-02 4.603921e-03
#> Locus_1461 0.000000e+00 0.000000e+00
#> Locus_1463 0.000000e+00 0.000000e+00
#> Locus_1468 2.316966e-02 -6.280326e-02
#> Locus_1470 -2.802146e-01 -7.632751e-02
#> Locus_1477 -5.058271e-03 7.012146e-02
#> Locus_1479 0.000000e+00 0.000000e+00
#> Locus_1482 -4.806203e-03 2.400778e-04
#> Locus_1485 3.219924e-02 1.322384e-02
#> Locus_1486 4.735535e-04 -1.926716e-02
#> Locus_1487 -2.364744e-02 -4.232024e-03
#> Locus_1488 -2.748919e-03 -1.081067e-02
#> Locus_1489 -1.401990e-02 -4.087007e-03
#> Locus_1490 -8.579472e-02 -1.276616e-02
#> Locus_1494 -2.491167e-02 -1.030996e-02
#> Locus_1496 -2.635690e-02 1.657123e-02
#> Locus_1500 2.916051e-02 2.516013e-02
#> Locus_1502 -1.002254e-02 2.261653e-02
#> Locus_1506 -2.252323e-02 -4.387478e-03
#> Locus_1508 -2.263976e-03 5.034704e-02
#> Locus_1509 4.437589e-04 2.172305e-02
#> Locus_1514 -1.911142e-02 -6.517701e-03
#> Locus_1515 -3.674862e-03 -2.556976e-02
#> Locus_1517 1.454252e-02 5.770477e-02
#> Locus_1520 -4.341493e-02 -7.133898e-03
#> Locus_1523 1.583812e-02 2.973030e-02
#> Locus_1524 -8.800870e-02 -4.591114e-02
#> Locus_1529 -2.052159e-02 -2.463376e-02
#> Locus_1534 -8.238139e-04 -3.417339e-03
#> Locus_1536 8.303560e-02 -1.241347e-01
#> Locus_1540 3.961032e-02 1.118002e-01
#> Locus_1545 -2.479348e-01 2.661481e-04
#> Locus_1549 1.779301e-02 2.923775e-02
#> Locus_1551 1.776998e-02 1.548991e-02
#> Locus_1554 -6.311614e-02 -5.868570e-02
#> Locus_1555 2.075684e-02 1.068474e-02
#> Locus_1561 -5.981247e-02 -6.485644e-02
#> Locus_1566 -7.099260e-02 -1.607517e-02
#> Locus_1572 0.000000e+00 0.000000e+00
#> Locus_1573 3.575300e-02 -3.228834e-02
#> Locus_1576 1.539051e-02 9.444929e-02
#> Locus_1586 8.242223e-02 -6.489884e-02
#> Locus_1587 -1.602648e-02 -5.123493e-03
#> Locus_1590 4.557072e-02 -3.562205e-02
#> Locus_1591 0.000000e+00 0.000000e+00
#> Locus_1596 0.000000e+00 0.000000e+00
#> Locus_1597 1.665479e-03 -3.167868e-02
#> Locus_1598 -4.034889e-02 -5.111902e-03
#> Locus_1601 2.856831e-02 9.291298e-04
#> Locus_1602 1.192371e-02 -4.191330e-03
#> Locus_1603 -1.326819e-02 -3.230225e-03
#> Locus_1604 -2.534819e-02 -1.119625e-02
#> Locus_1611 -1.191773e-02 5.754246e-02
#> Locus_1612 -1.522939e-02 4.816938e-04
#> Locus_1614 6.851495e-02 -5.134228e-02
#> Locus_1618 -3.278440e-02 -3.458712e-03
#> Locus_1619 -2.866217e-01 -1.034332e-01
#> Locus_1620 -5.269843e-02 -1.280566e-03
#> Locus_1621 0.000000e+00 0.000000e+00
#> Locus_1624 -1.845626e-02 -7.876140e-03
#> Locus_1626 1.177181e-02 -3.511073e-03
#> Locus_1632 -2.604218e-02 -1.298977e-02
#> Locus_1633 -9.227902e-03 2.353569e-03
#> Locus_1635 3.245404e-02 3.952705e-02
#> Locus_1636 2.538564e-02 -5.770307e-03
#> Locus_1641 0.000000e+00 0.000000e+00
#> Locus_1644 4.113179e-03 -3.490869e-02
#> Locus_1645 0.000000e+00 0.000000e+00
#> Locus_1646 1.289230e-01 -1.295463e-01
#> Locus_1647 -1.214118e-02 -4.285337e-03
#> Locus_1648 -4.956771e-02 2.430989e-02
#> Locus_1649 -4.309913e-02 -6.074020e-02
#> Locus_1652 -3.577389e-02 -3.899787e-02
#> Locus_1656 -1.942154e-02 -2.858330e-02
#> Locus_1658 -8.387120e-02 -5.592310e-03
#> Locus_1660 7.366474e-02 2.101135e-02
#> Locus_1661 -2.848623e-02 -4.155840e-02
#> Locus_1664 3.238494e-02 1.191680e-01
#> Locus_1665 0.000000e+00 0.000000e+00
#> Locus_1670 -6.998822e-03 -1.455811e-02
#> Locus_1681 4.946809e-03 -3.013720e-03
#> Locus_1682 5.338922e-02 3.596175e-03
#> Locus_1687 0.000000e+00 0.000000e+00
#> Locus_1690 0.000000e+00 0.000000e+00
#> Locus_1691 -2.207434e-02 -1.414645e-02
#> Locus_1692 1.067120e-02 1.878199e-02
#> Locus_1693 -5.122891e-02 1.492803e-02
#> Locus_1694 1.280674e-01 -1.263574e-01
#> Locus_1697 -5.531077e-02 -4.109726e-02
#> Locus_1699 -3.553527e-02 -4.454441e-02
#> Locus_1702 -2.556683e-02 -8.951518e-03
#> Locus_1703 3.141458e-02 -6.373600e-04
#> Locus_1708 7.779606e-03 -1.033632e-02
#> Locus_1713 -2.760888e-02 5.005098e-03
#> Locus_1715 0.000000e+00 0.000000e+00
#> Locus_1724 2.214026e-01 -1.541909e-01
#> Locus_1728 5.427338e-02 -5.222101e-02
#> Locus_1730 -5.424100e-02 6.149801e-03
#> Locus_1734 4.677777e-02 -3.420810e-02
#> Locus_1737 1.038433e-01 -1.037806e-01
#> Locus_1742 1.456158e-01 -1.264038e-01
#> Locus_1745 -1.798440e-01 1.105715e-03
#> Locus_1748 0.000000e+00 0.000000e+00
#> Locus_1750 -2.131764e-02 -5.136271e-02
#> Locus_1751 0.000000e+00 0.000000e+00
#> Locus_1755 -1.692398e-02 -6.940605e-03
#> Locus_1756 1.963806e-02 5.226529e-02
#> Locus_1757 -2.649095e-02 -1.932925e-04
#> Locus_1774 0.000000e+00 0.000000e+00
#> Locus_1776 1.108671e-03 -1.164717e-02
#> Locus_1777 -2.526596e-02 -4.986645e-03
#> Locus_1778 1.902275e-02 1.645254e-02
#> Locus_1783 0.000000e+00 0.000000e+00
#> Locus_1784 -1.427037e-01 -9.736977e-02
#> Locus_1786 -4.828837e-02 2.369225e-02
#> Locus_1787 3.233554e-02 8.694253e-02
#> Locus_1797 0.000000e+00 0.000000e+00
#> Locus_1799 0.000000e+00 0.000000e+00
#> Locus_1801 1.634857e-01 -1.346456e-01
#> Locus_1804 0.000000e+00 0.000000e+00
#> Locus_1805 -9.902291e-02 -5.977031e-02
#> Locus_1807 -1.138808e-01 -6.451638e-03
#> Locus_1808 3.628470e-02 1.153460e-01
#> Locus_1811 -2.097036e-02 -1.183728e-02
#> Locus_1813 0.000000e+00 0.000000e+00
#> Locus_1816 1.454252e-02 5.770477e-02
#> Locus_1822 3.388147e-02 1.458717e-02
#> Locus_1823 -5.582242e-02 -5.685403e-02
#> Locus_1826 0.000000e+00 0.000000e+00
#> Locus_1828 1.186213e-02 3.716561e-02
#> Locus_1833 2.333298e-01 -1.995079e-01
#> Locus_1839 5.300246e-03 3.315879e-02
#> Locus_1840 -1.424340e-01 -1.935386e-02
#> Locus_1841 4.450819e-02 2.497996e-02
#> Locus_1842 -5.398817e-02 -1.021377e-01
#> Locus_1843 -7.656609e-03 -8.015437e-03
#> Locus_1844 1.409179e-01 -1.914383e-01
#> Locus_1846 1.195318e-02 -1.337919e-02
#> Locus_1848 4.450423e-02 -4.148779e-03
#> Locus_1849 -1.402806e-01 2.863952e-02
#> Locus_1852 -1.331332e-02 -1.715015e-02
#> Locus_1853 -3.851305e-02 -6.026797e-02
#> Locus_1856 -6.998822e-03 -1.455811e-02
#> Locus_1861 -1.406596e-02 -7.268130e-03
#> Locus_1863 1.751288e-02 -1.961781e-02
#> Locus_1868 -1.979319e-02 -8.150311e-03
#> Locus_1870 1.192371e-02 -4.191330e-03
#> Locus_1871 -1.794956e-02 5.329133e-03
#> Locus_1872 1.037194e-01 -9.156725e-02
#> Locus_1873 -1.209717e-01 -2.737199e-02
#> Locus_1881 -2.163112e-01 1.565425e-01
#> Locus_1882 1.824012e-02 -1.267053e-02
#> Locus_1887 0.000000e+00 0.000000e+00
#> Locus_1894 -3.665088e-02 -4.174803e-02
#> Locus_1896 -1.326819e-02 -3.230225e-03
#> Locus_1908 -4.011342e-02 5.221884e-02
#> Locus_1911 9.361608e-02 -1.032126e-01
#> Locus_1913 -3.108921e-02 -1.639211e-04
#> Locus_1919 -1.522939e-02 4.816938e-04
#> Locus_1921 -3.865791e-03 -2.801275e-02
#> Locus_1932 0.000000e+00 0.000000e+00
#> Locus_1933 -9.956106e-03 4.527616e-03
#> Locus_1935 -6.017982e-02 -2.603701e-03
#> Locus_1939 -1.615633e-02 9.641833e-03
#> Locus_1941 -3.226661e-02 2.246479e-02
#> Locus_1942 -7.564493e-03 -1.653190e-03
#> Locus_1945 -2.854487e-02 -3.745220e-02
#> Locus_1948 -2.084851e-02 -2.982245e-02
#> Locus_1949 1.755713e-01 -1.848189e-01
#> Locus_1950 -3.721768e-02 2.344299e-02
#> Locus_1951 -8.703503e-03 -1.003190e-02
#> Locus_1952 -6.314747e-03 -3.688706e-03
#> Locus_1953 4.898173e-04 2.490417e-02
#> Locus_1958 2.754838e-02 5.443612e-02
#> Locus_1961 -9.465606e-02 2.984373e-02
#> Locus_1962 -3.199252e-01 -2.566994e-01
#> Locus_1965 6.278894e-03 6.545809e-02
#> Locus_1967 -1.203596e-01 -5.207401e-03
#> Locus_1968 -3.226661e-02 2.246479e-02
#> Locus_1971 2.272747e-02 2.284879e-02
#> Locus_1974 6.851506e-03 4.167267e-02
#> Locus_1977 5.721065e-02 -5.110564e-02
#> Locus_1982 1.987298e-03 -3.545959e-02
#> Locus_1983 -2.367500e-02 2.386865e-02
#> Locus_1984 1.402562e-02 1.424962e-02
#> Locus_1988 -5.073122e-02 5.016413e-03
#> Locus_1990 -4.985866e-03 -2.981695e-02
#> Locus_1991 0.000000e+00 0.000000e+00
#> Locus_1993 0.000000e+00 0.000000e+00
#> Locus_2003 -4.326920e-02 -7.692321e-03
#> Locus_2007 -1.203596e-01 -5.207401e-03
#> Locus_2014 2.487013e-02 -2.728533e-02
#> Locus_2016 4.609125e-02 -3.862840e-02
#> Locus_2017 1.371056e-02 5.985850e-02
#> Locus_2019 -1.101665e-02 1.177710e-02
#> Locus_2020 2.173104e-02 1.364621e-02
#> Locus_2021 -6.634403e-03 4.526125e-02
#> Locus_2027 -2.649095e-02 -1.932925e-04
#> Locus_2030 0.000000e+00 0.000000e+00
#> Locus_2033 0.000000e+00 0.000000e+00
#> Locus_2036 2.065680e-02 4.233881e-03
#> Locus_2041 3.219924e-02 1.322384e-02
#> Locus_2045 2.792527e-01 -2.426175e-01
#> Locus_2048 0.000000e+00 0.000000e+00
#> Locus_2050 -3.456662e-02 -7.355444e-02
#> Locus_2053 1.487129e-01 -1.272075e-01
#> Locus_2055 2.501375e-02 -1.091628e-01
#> Locus_2060 1.421627e-03 -2.255467e-02
#> Locus_2063 -2.098904e-02 1.859855e-03
#> Locus_2064 -7.127852e-02 -2.323801e-02
#> Locus_2068 0.000000e+00 0.000000e+00
#> Locus_2069 1.172875e-01 -1.119856e-01
#> Locus_2080 3.061729e-02 -7.677460e-03
#> Locus_2085 -1.345912e-02 -5.673221e-03
#> Locus_2086 -3.884309e-02 -5.716661e-02
#> Locus_2087 -1.942154e-02 -2.858330e-02
#> Locus_2091 -7.979633e-03 -1.792373e-03
#> Locus_2094 -4.079549e-02 -8.518353e-03
#> Locus_2102 -1.076587e-01 4.557252e-03
#> Locus_2106 0.000000e+00 0.000000e+00
#> Locus_2108 9.361608e-02 -1.032126e-01
#> Locus_2109 2.929516e-02 -3.198850e-02
#> Locus_2113 7.886378e-03 3.031139e-02
#> Locus_2115 -1.491740e-02 -5.904120e-03
#> Locus_2120 4.485178e-02 1.287521e-01
#> Locus_2128 -2.122974e-02 -2.349075e-02
#> Locus_2131 -2.518424e-02 8.484421e-03
#> Locus_2133 -1.920799e-01 1.668867e-03
#> Locus_2135 -4.806203e-03 2.400778e-04
#> Locus_2136 -1.062492e-01 9.950661e-03
#> Locus_2137 7.150600e-02 -6.457668e-02
#> Locus_2138 -2.207434e-02 -1.414645e-02
#> Locus_2139 -2.974218e-02 3.156558e-03
#> Locus_2140 -4.217935e-02 6.246974e-03
#> Locus_2145 -1.405623e-01 7.647038e-03
#> Locus_2148 0.000000e+00 0.000000e+00
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#> Locus_2157 -4.914454e-02 1.870531e-02
#> Locus_2159 2.804581e-02 5.287906e-02
#> Locus_2162 -8.632350e-02 1.436886e-01
#> Locus_2165 -1.807440e-02 -2.990147e-03
#> Locus_2166 1.187185e-02 2.939789e-03
#> Locus_2168 0.000000e+00 0.000000e+00
#> Locus_2179 4.321975e-02 1.694936e-03
#> Locus_2180 7.500340e-02 1.540309e-01
#> Locus_2181 3.588492e-02 9.526876e-02
#> Locus_2183 -1.401297e-01 -1.208159e-01
#> Locus_2185 -1.195626e-01 7.548834e-03
#> Locus_2186 2.674103e-02 6.874678e-02
#> Locus_2188 -4.997132e-03 -2.202919e-03
#> Locus_2191 -1.760708e-02 -3.105680e-03
#> Locus_2203 2.779292e-02 7.767302e-02
#> Locus_2208 4.055779e-02 9.137685e-03
#> Locus_2211 -6.512210e-02 5.984420e-02
#> Locus_2213 2.836553e-02 5.388760e-02
#> Locus_2215 -1.570645e-01 -7.056792e-02
#> Locus_2216 4.542343e-03 7.494962e-03
#> Locus_2219 -2.094665e-01 -8.385121e-02
#> Locus_2223 -5.043190e-03 -5.384042e-03
#> Locus_2228 0.000000e+00 0.000000e+00
#> Locus_2229 -1.081279e-02 -1.347147e-02
#> Locus_2230 0.000000e+00 0.000000e+00
#> Locus_2231 -6.455412e-03 -2.433817e-03
#> Locus_2234 -4.605840e-05 -3.181123e-03
#> Locus_2236 -3.067916e-02 -2.299957e-02
#> Locus_2238 -2.012612e-02 -5.509299e-03
#> Locus_2243 -7.327197e-03 1.366801e-02
#> Locus_2246 -6.465891e-02 -8.112959e-02
#> Locus_2250 1.847914e-02 5.967111e-04
#> Locus_2251 8.150919e-03 -7.784364e-03
#> Locus_2254 -1.186295e-01 -1.131354e-01
#> Locus_2260 1.883746e-02 -3.312063e-04
#> Locus_2261 -4.213111e-02 -7.520763e-02
#> Locus_2267 1.116584e-02 -4.055216e-02
#> Locus_2268 -1.469460e-02 6.839504e-03
#> Locus_2271 0.000000e+00 0.000000e+00
#> Locus_2273 0.000000e+00 0.000000e+00
#> Locus_2274 3.456692e-02 1.090769e-02
#> Locus_2275 3.570579e-02 1.095654e-01
#> Locus_2277 -1.047425e-02 2.150001e-02
#> Locus_2282 3.613663e-03 -5.638294e-02
#> Locus_2287 5.424420e-02 2.575453e-01
#> Locus_2294 -9.902291e-02 -5.977031e-02
#> Locus_2298 2.650123e-03 1.657940e-02
#> Locus_2305 -1.101204e-01 1.765231e-02
#> Locus_2308 3.318339e-02 -1.060661e-02
#> Locus_2309 4.557072e-02 -3.562205e-02
#> Locus_2312 5.654678e-02 1.867285e-01
#> Locus_2318 2.441612e-02 9.721654e-02
#> Locus_2319 -5.998961e-04 1.052323e-01
#> Locus_2321 -4.345036e-02 1.297878e-01
#> Locus_2322 -1.641419e-01 -1.653327e-01
#> Locus_2323 -4.250321e-02 -3.858274e-02
#> Locus_2325 -6.038244e-02 -9.751742e-02
#> Locus_2326 1.624686e-02 4.300347e-02
#> Locus_2327 -1.678342e-01 -9.074333e-02
#> Locus_2333 0.000000e+00 0.000000e+00
#> Locus_2336 0.000000e+00 0.000000e+00
#> Locus_2338 2.706741e-01 -6.083377e-02
#> Locus_2339 -5.043190e-03 -5.384042e-03
#> Locus_2344 1.361317e-01 -9.939497e-02
#> Locus_2345 1.824012e-02 -1.267053e-02
#> Locus_2348 0.000000e+00 0.000000e+00
#> Locus_2349 -9.612406e-03 4.801555e-04
#> Locus_2354 0.000000e+00 0.000000e+00
#> Locus_2359 1.187765e-02 -7.372453e-03
#> Locus_2361 1.156782e-01 -6.823343e-02
#> Locus_2364 -5.061921e-02 -5.159866e-02
#> Locus_2365 0.000000e+00 0.000000e+00
#> Locus_2366 -3.956818e-02 2.233359e-02
#> Locus_2368 8.468780e-02 -2.507087e-02
#> Locus_2371 1.776998e-02 1.548991e-02
#> Locus_2375 -5.717011e-02 -2.319287e-02
#> Locus_2381 -2.084851e-02 -2.982245e-02
#> Locus_2388 1.728829e-02 3.445762e-03
#> Locus_2394 -6.555576e-02 6.134587e-02
#> Locus_2400 6.069948e-02 1.291462e-01
#> Locus_2402 -1.204659e-02 -1.740445e-02
#> Locus_2406 -3.385581e-02 -8.483177e-02
#> Locus_2413 2.946607e-02 -3.299700e-02
#> Locus_2414 2.065680e-02 4.233881e-03
#> Locus_2415 3.383181e-02 4.131872e-03
#> Locus_2421 -8.116886e-02 -7.438455e-04
#> Locus_2424 2.969369e-02 1.129858e-02
#> Locus_2425 1.136406e-01 2.098404e-01
#> Locus_2426 -1.501018e-01 -2.050843e-03
#> Locus_2431 1.780045e-01 -1.748843e-01
#> Locus_2432 1.902275e-02 1.645254e-02
#> Locus_2433 1.092642e-01 -9.102343e-02
#> Locus_2434 -1.203596e-01 -5.207401e-03
#> Locus_2435 1.751288e-02 -1.961781e-02
#> Locus_2436 8.107274e-02 7.957389e-02
#> Locus_2437 9.361608e-02 -1.032126e-01
#> Locus_2439 -1.209253e-02 1.647873e-02
#> Locus_2448 -3.530952e-01 -2.112150e-01
#> Locus_2449 -1.498400e-01 -6.524345e-02
#> Locus_2450 0.000000e+00 0.000000e+00
#> Locus_2451 -2.560748e-02 -3.051344e-02
#> Locus_2452 0.000000e+00 0.000000e+00
#> Locus_2457 -2.360781e-01 -1.040444e-01
#> Locus_2461 5.192165e-02 -5.189031e-02
#> Locus_2463 -4.747453e-02 -8.553593e-02
#> Locus_2466 0.000000e+00 0.000000e+00
#> Locus_2467 1.395749e-01 -1.099643e-01
#> Locus_2473 0.000000e+00 0.000000e+00
#> Locus_2475 0.000000e+00 0.000000e+00
#> Locus_2477 1.754564e-01 -2.849499e-01
#> Locus_2480 7.960927e-02 2.013198e-01
#> Locus_2494 0.000000e+00 0.000000e+00
#> Locus_2497 -2.696431e-02 -1.452757e-02
#> Locus_2501 -1.586168e-02 -5.933189e-02
#> Locus_2502 -1.512899e-02 -3.306380e-03
#> Locus_2507 0.000000e+00 0.000000e+00
#> Locus_2510 2.182252e-02 6.722494e-02
#> Locus_2512 -9.364744e-02 -4.975387e-02
#> Locus_2514 4.391991e-02 2.384165e-03
#> Locus_2517 5.026789e-02 1.212704e-01
#> Locus_2518 2.429577e-03 5.311137e-03
#> Locus_2519 -5.030183e-02 -6.849802e-02
#> Locus_2522 -8.194222e-02 -1.179068e-01
#> Locus_2526 1.388691e-01 -1.430162e-01
#> Locus_2529 -2.526511e-02 -1.586929e-02
#> Locus_2530 5.300794e-02 1.677747e-02
#> Locus_2532 3.732725e-02 9.599499e-02
#> Locus_2540 -3.067200e-02 -4.411856e-02
#> Locus_2542 2.369553e-02 -7.702403e-03
#> Locus_2546 0.000000e+00 0.000000e+00
#> Locus_2550 0.000000e+00 0.000000e+00
#> Locus_2555 9.507055e-02 -8.931805e-02
#> Locus_2559 2.507564e-01 -1.212282e-01
#> Locus_2561 -9.535240e-02 -8.109618e-02
#> Locus_2565 -1.728331e-02 -3.677722e-02
#> Locus_2566 -1.203596e-01 -5.207401e-03
#> Locus_2567 1.949692e-02 2.010477e-02
#> Locus_2572 9.114145e-02 -7.124409e-02
#> Locus_2573 5.397776e-04 -6.193152e-02
#> Locus_2577 2.868524e-02 5.489614e-02
#> Locus_2579 -9.679418e-02 -5.670254e-02
#> Locus_2581 -2.869638e-01 -1.333452e-01
#> Locus_2583 9.805834e-03 -5.663505e-02
#> Locus_2588 6.809962e-02 1.004182e-01
#> Locus_2589 1.110880e-01 -8.254670e-04
#> Locus_2592 -2.320935e-02 -6.444063e-03
#> Locus_2597 1.187185e-02 2.939789e-03
#> Locus_2598 2.470545e-02 9.278771e-02
#> Locus_2599 -1.942154e-02 -2.858330e-02
#> Locus_2601 0.000000e+00 0.000000e+00
#> Locus_2602 9.369089e-02 -7.644670e-02
#> Locus_2611 1.876365e-02 3.737238e-02
#> Locus_2616 0.000000e+00 0.000000e+00
#> Locus_2617 3.243660e-03 5.119033e-02
#> Locus_2619 -2.526596e-02 -4.986645e-03
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#> Locus_2629 0.000000e+00 0.000000e+00
#> Locus_2632 -7.200703e-02 5.890148e-02
#> Locus_2634 -3.518097e-02 1.925569e-02
#> Locus_2636 1.177619e-02 1.108190e-02
#> Locus_2639 -2.077013e-02 1.396465e-02
#> Locus_2644 -5.433458e-04 -1.934368e-02
#> Locus_2647 -7.937889e-02 1.895922e-02
#> Locus_2648 -1.256162e-02 -3.856109e-03
#> Locus_2652 0.000000e+00 0.000000e+00
#> Locus_2655 -8.386804e-03 -2.215834e-02
#> Locus_2656 0.000000e+00 0.000000e+00
#> Locus_2657 7.561769e-02 -1.000863e-01
#> Locus_2660 -3.759060e-02 -3.218634e-03
#> Locus_2662 -4.242533e-02 1.353554e-01
#> Locus_2665 -1.008057e-01 -2.230156e-02
#> Locus_2668 7.477621e-02 1.070518e-02
#> Locus_2669 -2.320670e-01 -3.838491e-02
#> Locus_2670 -1.839988e-01 -1.217248e-02
#> Locus_2671 -1.942154e-02 -2.858330e-02
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#> Locus_2685 -1.937693e-01 -1.359284e-01
#> Locus_2687 5.196119e-04 -1.608604e-02
#> Locus_2696 -5.161050e-02 -7.305642e-03
#> Locus_2698 7.169062e-02 1.734777e-01
#> Locus_2699 -1.501018e-01 -2.050843e-03
#> Locus_2702 8.792872e-03 1.382708e-03
#> Locus_2703 -8.808509e-02 -1.421147e-01
#> Locus_2705 -1.154849e-02 1.703483e-01
#> Locus_2712 -2.079465e-02 -6.415608e-03
#> Locus_2718 3.409861e-03 -5.305134e-04
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#> Locus_2725 -1.518209e-02 1.555225e-02
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#> Locus_2731 -7.960136e-02 -3.118700e-02
#> Locus_2733 4.526481e-02 -6.246609e-02
#> Locus_2736 1.925944e-02 -4.833948e-02
#> Locus_2738 2.218489e-02 -1.187002e-02
#> Locus_2739 -2.173019e-02 -6.700528e-03
#> Locus_2753 -3.813073e-02 -6.673156e-03
#> Locus_2755 -1.024711e-02 -2.637639e-02
#> Locus_2758 9.192580e-02 2.386635e-01
#> Locus_2760 2.939009e-02 -3.123490e-02
#> Locus_2761 1.902275e-02 1.645254e-02
#> Locus_2764 2.218489e-02 -1.187002e-02
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#> Locus_2767 7.202871e-03 1.406993e-01
#> Locus_2768 3.625209e-04 1.563421e-02
#> Locus_2770 -1.196642e-01 3.709416e-03
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#> Locus_2776 5.006044e-02 -1.681623e-02
#> Locus_2789 -9.443531e-02 -2.091154e-02
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#> Locus_2796 9.778126e-03 -1.614083e-02
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#> Locus_2809 -2.974218e-02 3.156558e-03
#> Locus_2810 -3.088423e-02 -4.617343e-02
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#> Locus_2814 6.074609e-03 9.529356e-04
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#> Locus_2878 6.619645e-03 -1.396275e-02
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#> Locus_2938 4.544143e-02 -6.027241e-02
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#> Locus_2991 9.817152e-03 1.380477e-02
#> Locus_2993 1.458980e-01 -1.417937e-01
#> Locus_2994 -3.389695e-02 -1.012164e-02
#> Locus_2995 3.804550e-02 3.290508e-02
#> Locus_3005 1.216966e-02 3.079693e-02
#> Locus_3009 5.380650e-02 -1.857759e-02
#> Locus_3010 -8.571832e-02 -6.568890e-02
#> Locus_3012 -1.180919e-01 7.672256e-03
#> Locus_3013 0.000000e+00 0.000000e+00
#> Locus_3023 1.395216e-02 7.581178e-02
#> Locus_3029 -2.690721e-02 -1.098982e-02
#> Locus_3030 3.494879e-02 1.308630e-01
#> Locus_3032 9.114145e-02 -7.124409e-02
#> Locus_3042 -2.110741e-02 1.062004e-02
#> Locus_3043 1.081444e-02 -8.522131e-04
#> Locus_3046 2.190958e-02 5.196513e-03
#> Locus_3047 0.000000e+00 0.000000e+00
#> Locus_3051 -2.248515e-02 6.814000e-03
#> Locus_3054 -1.257517e-02 3.259277e-02
#> Locus_3056 2.150561e-02 1.120833e-01
#> Locus_3058 -2.526987e-01 2.640794e-02
#> Locus_3059 -4.169701e-02 -5.964490e-02
#> Locus_3060 1.882605e-02 4.418972e-02
#> Locus_3061 -9.882070e-03 -1.888542e-02
#> Locus_3062 6.763475e-02 1.203133e-01
#> Locus_3063 8.572116e-03 6.621892e-02
#> Locus_3064 2.916051e-02 2.516013e-02
#> Locus_3069 1.396835e-02 -3.295743e-02
#> Locus_3078 0.000000e+00 0.000000e+00
#> Locus_3079 -2.110741e-02 1.062004e-02
#> Locus_3081 -1.401990e-02 -4.087007e-03
#> Locus_3084 1.221208e-02 -6.277075e-04
#> Locus_3087 -1.373801e-01 -1.785767e-02
#> Locus_3089 4.919457e-02 -1.709042e-02
#> Locus_3090 -1.771697e-02 -3.108803e-03
#> Locus_3091 -2.029309e-02 1.674941e-01
#> Locus_3092 2.752807e-02 -6.854905e-02
#> Locus_3093 -1.345423e-02 -1.699192e-02
#> Locus_3097 7.108136e-04 -1.127733e-02
#> Locus_3098 -4.739560e-02 1.915841e-02
#> Locus_3102 0.000000e+00 0.000000e+00
#> Locus_3116 -8.992200e-02 5.528576e-04
#> Locus_3122 -3.059623e-01 1.733242e-01
#> Locus_3126 3.322640e-02 7.922560e-02
#> Locus_3128 -3.454803e-02 -8.127531e-04
#> Locus_3129 1.447217e-02 3.884035e-02
#> Locus_3133 -4.898173e-04 -2.490417e-02
#> Locus_3138 -2.158700e-02 -9.424665e-03
#> Locus_3140 0.000000e+00 0.000000e+00
#> Locus_3143 -1.525107e-01 -2.092317e-01
#> Locus_3145 0.000000e+00 0.000000e+00
#> Locus_3147 -1.031519e-01 -1.275645e-01
#> Locus_3148 1.114456e-01 -7.436556e-02
#> Locus_3154 5.780417e-04 1.224815e-02
#> Locus_3155 5.575659e-03 2.422147e-02
#> Locus_3160 1.855222e-01 -1.541265e-01
#> Locus_3161 -2.137863e-02 -6.040353e-02
#> Locus_3164 -1.731957e-02 4.162616e-02
#> Locus_3165 -1.817079e-02 -1.463260e-02
#> Locus_3167 -1.463319e-02 -2.019782e-02
#> Locus_3168 -1.525405e-02 9.430354e-03
#> Locus_3169 1.285135e-01 -1.323120e-01
#> Locus_3171 3.979048e-02 3.598132e-02
#> Locus_3175 -2.894082e-02 -3.055640e-02
#> Locus_3177 -2.929695e-01 -1.619309e-01
#> Locus_3179 1.409590e-01 -1.207585e-01
#> Locus_3181 3.106706e-03 1.712880e-02
#> Locus_3182 0.000000e+00 0.000000e+00
#> Locus_3188 -2.077471e-01 2.069357e-02
#> Locus_3190 0.000000e+00 0.000000e+00
#> Locus_3191 2.887690e-03 5.619626e-02
#> Locus_3192 1.742896e-02 2.190873e-03
#> Locus_3197 1.341604e-01 -6.537660e-02
#> Locus_3198 4.525299e-02 -3.980362e-02
#>
#> $sites
#> RDA1 RDA2 RDA3 PC1 PC2 PC3
#> ALT3 0.12259277 -2.03133124 -1.21538101 -1.45893312 -5.196493950 -0.22482810
#> BAR360 -0.10566063 0.40074788 1.80638271 3.08484692 -0.653189728 -0.14275205
#> BLL5 1.05001767 0.27503087 -1.85047689 -0.37364740 0.061378362 -0.97379152
#> BNT5 -3.08724981 -1.02639189 -1.11624992 -0.32196431 0.957826272 -0.51248485
#> BOF1 1.47216292 -1.10167029 0.25328114 -0.23873838 0.851335568 0.92449342
#> BOT3 0.81329232 1.47615245 -3.28470108 -0.95110772 -0.838349738 -0.96385781
#> CAR1 0.02419516 0.99797233 -0.16571641 -0.32517766 0.816402141 -0.15797135
#> CTR7 2.28856230 -2.48550913 -1.04473019 -0.69966660 0.921452039 1.62181457
#> CYR26 -0.21664805 0.85222570 -0.69242487 -0.69176836 0.031303481 1.35000535
#> FCT13 -0.05724046 1.66926170 4.21520334 2.89130967 -0.431499557 -0.19022081
#> HAY2 0.50081099 2.23498406 -2.56257017 -1.02251946 -1.315049404 0.04159399
#> HHR2 2.52273585 -3.00625328 -0.32689207 -0.79235741 0.774283688 2.89126436
#> HNT18 -3.45843529 -0.94121492 0.25433822 -0.51536579 1.575023982 -1.09409295
#> HOP1 2.10034811 -0.34925722 -0.26452947 -1.09452878 -1.492404225 -3.17569229
#> INK1 1.91409104 -1.87959312 1.01801433 -0.56094096 0.607761158 -0.55549629
#> INK6 1.62441083 -0.89667572 0.80876895 -0.35572941 0.769880312 -0.90028836
#> KBG6 0.44370116 -0.08071891 1.73375321 -0.82916349 -0.177585139 1.55701304
#> KRV3 0.03101360 0.32013109 2.15004275 -1.11474307 0.021147726 1.07523066
#> LBH40 -0.16538441 2.16474751 0.52801213 0.87615663 -0.340772346 1.14727874
#> LCV19 -3.31928114 -1.38467453 -1.13915781 -0.58133668 1.032150699 -1.15528558
#> LFV10 0.45048212 -1.99469509 -3.01758322 -0.18058479 0.036611579 1.58643411
#> LTF1 -0.56332964 -1.41319468 -0.40444124 -1.01455489 -2.568220862 0.27256705
#> LTF6 -0.80533934 -1.26200321 -0.40595923 -0.82166787 -1.812391612 0.16059745
#> MLG2 -0.19882166 0.24542728 0.97102328 2.19529163 -0.415012950 0.02073058
#> MLG5 -0.16359948 0.27579424 0.40464827 2.78533793 -0.730688253 -0.29965871
#> MNR2 0.29989192 1.34343687 0.08327683 -0.85296324 -0.896744546 0.60291973
#> MRO3 1.08828774 -0.85124054 -0.85893929 -0.43982456 0.228836668 1.43162166
#> OLA14 -2.81168154 -0.91474657 -0.05525087 -0.62705876 1.512227726 -1.69398713
#> OLA9 -3.75268081 -0.65996391 0.06256908 -0.28133154 2.529620489 -2.76218883
#> POT1 1.24145606 -2.42380913 -3.15924222 -0.53307464 -0.232104652 2.76100504
#> PRU1 -1.05358457 1.60027065 1.51580134 0.10208565 1.016489574 -0.30317924
#> PRU6 -0.61460509 2.49853741 1.78494576 0.38406109 1.029605881 -0.36191898
#> PT8 0.50479265 1.69127823 0.52456816 -1.16343339 -0.972429139 -0.05828469
#> REY5 -0.55582361 1.85425742 -0.76456682 -0.37383101 0.871101814 -0.15091350
#> RPS38 -3.37961550 -1.20914098 -1.05406461 -0.49298007 1.981221332 -1.35743366
#> RPS9 -3.55780994 -1.10616773 -0.50205282 -0.07694763 1.953785285 -1.71851026
#> SBR10 -0.53162025 1.84734378 -1.63544918 -0.49265297 1.025127868 0.25384918
#> SKY3 1.08632380 1.49006553 -2.99538153 -0.90793440 -0.933677978 -1.16325378
#> SKY5 1.03751234 1.47238216 -4.09914131 -0.91420827 -0.866578877 -1.41189823
#> SRM1 -0.28353104 0.28046598 2.57546667 2.63982023 -0.557421246 -0.21015874
#> SRM7 -0.11572481 0.58300041 2.62416301 2.89247927 -1.088862339 -0.07795291
#> SUT7 0.93108534 0.15987771 0.78309974 -1.16573237 -1.114590875 -1.03459518
#> TTC1 1.25775103 -1.33162033 -0.07392339 -0.39358363 0.768208383 1.26233187
#> TTC2 1.41991380 -1.53038614 0.42293394 -0.42887838 0.767394885 1.52249097
#> WAW1 1.28052589 -1.31959679 -0.07316748 -0.53515004 0.823728341 0.83740665
#> WDC1 0.89115235 -0.29289209 1.16692957 -0.62558934 0.569387961 0.56778677
#> WDC2 2.17770630 -1.92881436 1.38136902 -1.00727193 0.481455052 2.09151079
#> WHR2 1.95631786 0.02594610 -2.51822161 -1.07671648 -1.677039590 -2.46815241
#> WTW14 -0.27082248 0.58948598 1.89404890 2.42733174 0.048845311 -1.11433241
#> WTW3 0.14670024 0.28258808 2.55064742 2.36324932 -0.003977117 -0.27468822
#> YUC3 -0.32626611 0.46764343 4.34101858 2.31054970 -1.391113403 1.10725516
#> ZUM18 -0.78452876 3.41895826 -0.45577769 0.73256113 0.875390080 0.75189524
#> ZUM20 -0.49854974 2.90354870 -0.11831398 0.64857788 0.767213868 0.66877247
#>
#> $constraints
#> RDA1 RDA2 RDA3 PC1 PC2
#> ALT3 -1.86406684 -1.562324125 -0.26787615 -1.45893312 -5.196493950
#> BAR360 0.16413907 -0.964704173 -0.52878026 3.08484692 -0.653189728
#> BLL5 1.84297909 0.623915182 -2.87800799 -0.37364740 0.061378362
#> BNT5 -2.29910391 -1.271131510 -0.62271896 -0.32196431 0.957826272
#> BOF1 1.47221056 -1.068028205 0.16440937 -0.23873838 0.851335568
#> BOT3 0.91416246 1.354769305 -2.00633548 -0.95110772 -0.838349738
#> CAR1 0.52403255 0.394028192 -0.23953636 -0.32517766 0.816402141
#> CTR7 1.83678679 -1.284927411 -0.48594122 -0.69966660 0.921452039
#> CYR26 -1.33848752 2.723014046 -1.77642798 -0.69176836 0.031303481
#> FCT13 0.59192979 1.456740039 1.04679888 2.89130967 -0.431499557
#> HAY2 -0.41256612 1.878230692 -0.86041438 -1.02251946 -1.315049404
#> HHR2 1.21850122 -0.976017957 0.21173301 -0.79235741 0.774283688
#> HNT18 -2.26191241 -0.644022973 1.20052072 -0.51536579 1.575023982
#> HOP1 2.40421619 -0.185225885 0.87126280 -1.09452878 -1.492404225
#> INK1 2.31683965 -1.645686840 0.80513739 -0.56094096 0.607761158
#> INK6 2.31683965 -1.645686840 0.80513739 -0.35572941 0.769880312
#> KBG6 -0.70417325 0.956840666 2.37693611 -0.82916349 -0.177585139
#> KRV3 -0.78313902 1.078009565 3.20040778 -1.11474307 0.021147726
#> LBH40 -0.87120689 2.781732912 -0.37921517 0.87615663 -0.340772346
#> LCV19 -2.48830470 -1.175225591 -0.84336361 -0.58133668 1.032150699
#> LFV10 -0.66647505 -2.059887617 -2.70285056 -0.18058479 0.036611579
#> LTF1 -1.71570698 -1.280437412 0.44814330 -1.01455489 -2.568220862
#> LTF6 -1.71570698 -1.280437412 0.44814330 -0.82166787 -1.812391612
#> MLG2 0.06680820 -0.669293489 -0.72042464 2.19529163 -0.415012950
#> MLG5 0.06680820 -0.669293489 -0.72042464 2.78533793 -0.730688253
#> MNR2 -0.51998733 0.907902625 1.60141074 -0.85296324 -0.896744546
#> MRO3 0.32982582 -0.554791505 -1.02741866 -0.43982456 0.228836668
#> OLA14 -1.29028513 -0.395800048 0.30875182 -0.62705876 1.512227726
#> OLA9 -1.29028513 -0.395800048 0.30875182 -0.28133154 2.529620489
#> POT1 -0.80621474 -1.734231592 -3.01188642 -0.53307464 -0.232104652
#> PRU1 -0.13177772 0.953460230 1.16054379 0.10208565 1.016489574
#> PRU6 -0.13177772 0.953460230 1.16054379 0.38406109 1.029605881
#> PT8 -0.50268351 0.780606765 1.48904785 -1.16343339 -0.972429139
#> REY5 0.06737857 0.733336327 -0.55002217 -0.37383101 0.871101814
#> RPS38 -1.91015065 -0.976675106 -0.42713141 -0.49298007 1.981221332
#> RPS9 -1.91015065 -0.976675106 -0.42713141 -0.07694763 1.953785285
#> SBR10 -0.17716743 2.595590096 -1.10994635 -0.49265297 1.025127868
#> SKY3 1.05851774 0.722407799 -1.02392761 -0.90793440 -0.933677978
#> SKY5 1.05851774 0.722407799 -1.02392761 -0.91420827 -0.866578877
#> SRM1 -0.23609803 -0.477581684 0.24042568 2.63982023 -0.557421246
#> SRM7 -0.23609803 -0.477581684 0.24042568 2.89247927 -1.088862339
#> SUT7 0.34550029 0.488582343 1.83993611 -1.16573237 -1.114590875
#> TTC1 0.97234001 -1.065890882 0.03152674 -0.39358363 0.768208383
#> TTC2 0.97234001 -1.065890882 0.03152674 -0.42887838 0.767394885
#> WAW1 1.65002159 -0.971450647 0.36792840 -0.53515004 0.823728341
#> WDC1 1.00307508 -0.004432834 1.85014838 -0.62558934 0.569387961
#> WDC2 1.00307508 -0.004432834 1.85014838 -1.00727193 0.481455052
#> WHR2 2.13555573 0.803560444 -1.20938677 -1.07671648 -1.677039590
#> WTW14 0.70904126 -0.413406849 0.48708744 2.42733174 0.048845311
#> WTW3 0.70904126 -0.413406849 0.48708744 2.36324932 -0.003977117
#> YUC3 -0.94621833 -0.018007196 1.71257372 2.31054970 -1.391113403
#> ZUM18 -0.27036978 2.709895708 -0.95169938 0.73256113 0.875390080
#> ZUM20 -0.27036978 2.709895708 -0.95169938 0.64857788 0.767213868
#> PC3
#> ALT3 -0.22482810
#> BAR360 -0.14275205
#> BLL5 -0.97379152
#> BNT5 -0.51248485
#> BOF1 0.92449342
#> BOT3 -0.96385781
#> CAR1 -0.15797135
#> CTR7 1.62181457
#> CYR26 1.35000535
#> FCT13 -0.19022081
#> HAY2 0.04159399
#> HHR2 2.89126436
#> HNT18 -1.09409295
#> HOP1 -3.17569229
#> INK1 -0.55549629
#> INK6 -0.90028836
#> KBG6 1.55701304
#> KRV3 1.07523066
#> LBH40 1.14727874
#> LCV19 -1.15528558
#> LFV10 1.58643411
#> LTF1 0.27256705
#> LTF6 0.16059745
#> MLG2 0.02073058
#> MLG5 -0.29965871
#> MNR2 0.60291973
#> MRO3 1.43162166
#> OLA14 -1.69398713
#> OLA9 -2.76218883
#> POT1 2.76100504
#> PRU1 -0.30317924
#> PRU6 -0.36191898
#> PT8 -0.05828469
#> REY5 -0.15091350
#> RPS38 -1.35743366
#> RPS9 -1.71851026
#> SBR10 0.25384918
#> SKY3 -1.16325378
#> SKY5 -1.41189823
#> SRM1 -0.21015874
#> SRM7 -0.07795291
#> SUT7 -1.03459518
#> TTC1 1.26233187
#> TTC2 1.52249097
#> WAW1 0.83740665
#> WDC1 0.56778677
#> WDC2 2.09151079
#> WHR2 -2.46815241
#> WTW14 -1.11433241
#> WTW3 -0.27468822
#> YUC3 1.10725516
#> ZUM18 0.75189524
#> ZUM20 0.66877247
#>
#> $biplot
#> RDA1 RDA2 RDA3 PC1 PC2 PC3
#> CA_rPCA1 -0.2047232 0.5349644 0.8196960 0 0 0
#> CA_rPCA2 0.5187643 0.8393239 -0.1625394 0 0 0
#> CA_rPCA3 0.5560921 -0.6173999 0.5563982 0 0 0
#>
#> $call
#> rda(formula = gen ~ CA_rPCA1 + CA_rPCA2 + CA_rPCA3, data = moddf)
#>
#> $tot.chi
#> [1] 143.389One of the most relevant statistics to present from an RDA model is the adjusted R2 value. The R2 value must be adjusted; if not, it is biased because it will always increase if independent variables are added (i.e., there is no penalization for adding independent variables that aren’t significantly affecting dependent variables within the model). The adjusted R2 value from the full (global) model can also help us determine the stopping point when we go on to do variable selection. As we can see, the unadjusted R2 is 0.098, while the adjusted value is 0.042. Be sure to always report adjusted R2 values.
RsquareAdj(mod_full)
#> $r.squared
#> [1] 0.0976769
#>
#> $adj.r.squared
#> [1] 0.04243263Run a simple RDA with variable selection
Now, let’s run an RDA model with variable selection by specifying the
"best" model within rda_run(). Variable
selection occurs using the ordiR2step() function in the
vegan package, and is a forward selection method that begins with a null
model and adds explanatory variables one at a time until genetic
variance is maximized. To perform forward selection, we need to specify
stopping criteria (i.e., when to stop adding more explanatory
variables). There are two primary ways to specify stopping criteria:
parameters involved in a permutation-based significance test (two
parameters: the limits of permutation p-values for adding
(Pin) or dropping (Pout) a term to the model,
and the number of permutations used in the estimation of adjusted R2
(R2permutations)), and whether to use the adjusted R2 value
as the stopping criterion (R2scope; only models with
adjusted R2 values lower than specified are accepted).
mod_best <- rda_run(gen, env,
model = "best",
Pin = 0.05,
R2permutations = 1000,
R2scope = T
)
#> Step: R2.adj= 0
#> Call: gen ~ 1
#>
#> R2.adjusted
#> <All variables> 0.0424326339
#> + CA_rPCA2 0.0196214014
#> + CA_rPCA3 0.0137427906
#> + CA_rPCA1 0.0009953656
#> <none> 0.0000000000
#>
#> Df AIC F Pr(>F)
#> + CA_rPCA2 1 264.09 2.0407 0.002 **
#> ---
#> Signif. codes: 0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
#>
#> Step: R2.adj= 0.0196214
#> Call: gen ~ CA_rPCA2
#>
#> R2.adjusted
#> <All variables> 0.04243263
#> + CA_rPCA3 0.03919420
#> + CA_rPCA1 0.01991617
#> <none> 0.01962140
#>
#> Df AIC F Pr(>F)
#> + CA_rPCA3 1 263.97 2.0389 0.006 **
#> ---
#> Signif. codes: 0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
#>
#> Step: R2.adj= 0.0391942
#> Call: gen ~ CA_rPCA2 + CA_rPCA3
#>
#> R2.adjusted
#> <All variables> 0.04243263
#> + CA_rPCA1 0.04243263
#> <none> 0.03919420
#>
#> Df AIC F Pr(>F)
#> + CA_rPCA1 1 264.72 1.1691 0.158Let’s look at our best model, and see how it compares to our full
model. There is also an additional element in the mod_best
object, which are the results from an ANOVA-like permutation test. When
we look at the call, we can now see that only environmental PCs 2 and 3
are considered in the model, meaning that PC1 is not considered to be
significantly associated with genetic variance. Importantly, our
adjusted R2 value is 0.039 (compared to 0.042 for the full model), which
tells us that the model with only two of the environmental variables
explains nearly as much as the RDA with all three.
mod_best$call
#> rda(formula = gen ~ CA_rPCA2 + CA_rPCA3, data = moddf)
mod_best$anova
#> R2.adj Df AIC F Pr(>F)
#> + CA_rPCA2 0.019621 1 264.09 2.0407 0.002 **
#> + CA_rPCA3 0.039194 1 263.97 2.0389 0.006 **
#> <All variables> 0.042433
#> ---
#> Signif. codes: 0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
RsquareAdj(mod_best)
#> $r.squared
#> [1] 0.07614827
#>
#> $adj.r.squared
#> [1] 0.0391942Following variable selection, we want to retain two environmental variables (CA_rPCA2 and CA_rPCA3) for all subsequent analyses.
Run a partial RDA with geography as a covariable
A partial RDA, as described at the beginning of the vignette, is when
covariables (or conditioning variables) are incorporated into the RDA.
Within rda_run(), to correct for geography, users can set
correctGEO = TRUE, in which case sampling coordinates
(coords) must also be provided. Let’s now run a partial
RDA, without any variable selection (the full model) and view the
results.
mod_pRDA_geo <- rda_run(gen, env, liz_coords,
model = "full",
correctGEO = TRUE,
correctPC = FALSE
)Let’s take a look at our partial RDA results. Note that now, the call
includes conditioning variables x and y, which correspond to our
latitude and longitude values. Conditioning variables are specified
within the rda() function using the
Condition() function. Finally, when we look at the summary
of the resulting object, we can see that along with total, constrained,
and unconstrained inertia (or variance) as with our simple RDA, there is
an additional row for conditioned inertia because our partial RDA
includes conditioning variables. Interestingly, the conditioned inertia
is higher than constrained inertia, suggesting that geography is more
associated with genetic variance than our environmental variables
are.
anova(mod_pRDA_geo)
#> Permutation test for rda under reduced model
#> Permutation: free
#> Number of permutations: 999
#>
#> Model: rda(formula = gen ~ CA_rPCA1 + CA_rPCA2 + CA_rPCA3 + Condition(x + y), data = moddf)
#> Df Variance F Pr(>F)
#> Model 3 11.306 1.5948 0.001 ***
#> Residual 47 111.068
#> ---
#> Signif. codes: 0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
RsquareAdj(mod_pRDA_geo) # 0.0305
#> $r.squared
#> [1] 0.07884828
#>
#> $adj.r.squared
#> [1] 0.03058243
head(summary(mod_pRDA_geo))
#> $species
#> RDA1 RDA2 RDA3 PC1
#> Locus_10 3.454904e-02 8.515902e-03 2.220254e-02 2.631181e-02
#> Locus_15 3.360753e-02 -8.041141e-02 -7.978641e-02 5.156274e-02
#> Locus_22 4.500162e-02 3.759728e-03 1.079207e-02 -8.322817e-02
#> Locus_28 -6.112164e-02 -6.035290e-02 -6.887637e-02 3.954925e-02
#> Locus_32 1.352317e-02 1.104829e-02 2.970871e-02 1.883250e-02
#> Locus_35 -5.728107e-32 -2.203291e-31 -4.551088e-32 5.383479e-17
#> Locus_37 6.860116e-32 2.237051e-31 -4.476787e-32 -1.034333e-16
#> Locus_61 3.934522e-02 -5.980984e-02 1.196345e-02 -2.918604e-01
#> Locus_71 2.986347e-02 1.121422e-02 6.979140e-03 -7.695562e-02
#> Locus_72 -1.545083e-32 -6.820479e-32 -8.929031e-32 -9.768433e-17
#> Locus_82 2.781229e-32 -1.190473e-31 4.858645e-32 -7.104360e-17
#> Locus_84 2.771565e-03 3.341097e-02 -9.762235e-03 -3.203079e-02
#> Locus_90 7.728174e-02 4.104709e-02 1.017414e-01 1.146894e-01
#> Locus_91 1.067950e-01 -2.660009e-03 -3.473157e-03 7.808693e-02
#> Locus_96 4.757011e-33 -7.420671e-33 1.061800e-32 4.854276e-17
#> Locus_104 -1.906922e-02 4.987332e-03 4.650002e-03 -1.838639e-02
#> Locus_122 4.147512e-02 -8.811375e-02 -2.024241e-02 -7.523186e-02
#> Locus_125 1.230328e-01 2.157433e-02 2.300501e-02 2.100864e-02
#> Locus_131 1.337085e-02 7.121972e-02 6.158163e-03 -9.101327e-02
#> Locus_133 2.119939e-33 -4.621468e-33 -8.071766e-33 6.943809e-17
#> Locus_138 4.310661e-02 -3.660331e-02 3.749070e-02 -5.413312e-02
#> Locus_154 -1.042319e-02 -4.510426e-02 3.495489e-02 -7.003381e-03
#> Locus_162 5.098057e-02 -6.680153e-02 5.172222e-02 1.243331e-01
#> Locus_166 1.715775e-01 4.493404e-03 2.854554e-02 1.339723e-01
#> Locus_177 3.063679e-33 2.979752e-33 -1.758679e-33 9.753449e-18
#> Locus_185 -2.167814e-33 -4.378181e-33 4.082131e-33 -5.921315e-17
#> Locus_191 5.372036e-03 1.256056e-01 -5.837993e-02 6.896932e-02
#> Locus_193 -1.166176e-01 9.064614e-03 4.727183e-03 -1.727680e-01
#> Locus_207 1.007193e-02 -3.686019e-02 -3.041322e-02 -6.883044e-03
#> Locus_211 3.238056e-02 -1.780411e-04 -1.350000e-03 2.729645e-02
#> Locus_213 -7.009199e-02 -1.481516e-02 -5.429505e-02 -1.242852e-01
#> Locus_214 6.278081e-02 -6.202426e-02 -7.686803e-02 5.993614e-02
#> Locus_222 -3.395328e-04 4.149163e-02 6.914401e-04 -2.014374e-02
#> Locus_227 3.303590e-02 -2.882048e-02 4.573652e-02 -8.954750e-02
#> Locus_242 1.373598e-02 5.634577e-03 -5.684737e-04 4.262278e-02
#> Locus_249 5.340500e-02 -1.159769e-02 -8.564976e-03 -8.682165e-02
#> Locus_254 -4.840189e-02 -2.614615e-02 1.224754e-02 1.523768e-02
#> Locus_256 -4.080687e-03 -8.463698e-02 1.063368e-01 -5.698050e-02
#> Locus_263 1.537921e-01 -2.433227e-02 -2.846107e-02 6.260198e-02
#> Locus_268 1.839054e-02 -3.900940e-02 -1.852116e-02 -2.842081e-02
#> Locus_279 -1.773042e-02 -1.318811e-02 -4.925725e-02 1.855175e-02
#> Locus_285 1.917728e-02 6.116014e-02 4.528122e-02 -9.130050e-02
#> Locus_287 -3.511454e-02 1.587835e-03 -2.467662e-02 6.559088e-02
#> Locus_292 -2.616863e-03 -2.690044e-02 -1.305445e-02 -3.371516e-02
#> Locus_294 -6.863725e-03 3.315755e-02 1.436881e-02 2.045180e-02
#> Locus_303 3.915318e-33 -1.367651e-33 6.549145e-33 -1.274005e-17
#> Locus_308 -3.817820e-02 -1.623095e-02 5.008129e-02 -1.343201e-02
#> Locus_312 -2.109527e-02 9.591928e-02 -5.827952e-02 5.473276e-02
#> Locus_313 2.022735e-33 1.485033e-33 -5.871657e-34 -3.727287e-17
#> Locus_314 3.874327e-02 3.735390e-02 1.130287e-02 5.117972e-02
#> Locus_317 6.619130e-03 4.528220e-02 6.134313e-03 -4.369634e-02
#> Locus_318 -6.176444e-02 -1.014601e-01 6.567454e-02 3.988799e-02
#> Locus_323 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_330 1.883137e-02 -2.083875e-02 -8.113092e-03 -4.331455e-02
#> Locus_331 8.964341e-03 1.670232e-02 -3.241430e-02 -9.393061e-03
#> Locus_333 1.351267e-02 -9.512096e-03 1.221009e-02 1.593435e-02
#> Locus_334 2.182253e-02 -6.310520e-03 7.133025e-03 2.836189e-02
#> Locus_336 -9.432033e-03 1.683788e-03 1.964340e-03 2.269822e-02
#> Locus_337 5.723982e-02 5.386876e-03 -1.202938e-02 3.963128e-02
#> Locus_339 1.915738e-01 -1.030641e-01 -8.566576e-02 -1.570152e-01
#> Locus_341 1.485354e-01 -3.253169e-02 -5.711847e-02 6.912568e-02
#> Locus_345 -1.625907e-02 4.877829e-02 -2.949307e-02 -2.028382e-02
#> Locus_350 1.342850e-01 1.823331e-02 1.747597e-02 -3.036977e-02
#> Locus_354 -1.500404e-02 2.131845e-02 -3.739861e-02 1.176975e-02
#> Locus_359 -1.176604e-01 -1.629749e-01 3.144508e-02 -1.764198e-01
#> Locus_365 -3.595863e-03 -1.709768e-02 -4.673547e-03 -4.631843e-03
#> Locus_366 4.677681e-02 -4.248955e-02 -1.323975e-03 -6.434199e-02
#> Locus_369 -4.874894e-02 -1.690320e-02 -1.954696e-02 2.211420e-02
#> Locus_372 1.058582e-03 2.926532e-02 -5.502049e-03 -3.710464e-02
#> Locus_389 -6.467634e-02 -8.561301e-02 2.191375e-02 8.539531e-03
#> Locus_391 4.234334e-02 -4.822490e-03 -7.344876e-03 3.424927e-02
#> Locus_392 9.433237e-02 -2.111857e-02 -2.348563e-02 8.458165e-02
#> Locus_393 1.220692e-02 -6.306359e-03 7.408401e-03 -4.944713e-02
#> Locus_394 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_396 4.244718e-02 1.577450e-02 1.615251e-02 -1.048488e-02
#> Locus_399 9.665049e-02 -1.164883e-02 -1.822048e-02 7.480192e-02
#> Locus_400 1.445987e-02 4.248963e-02 -3.923144e-02 -1.922970e-02
#> Locus_401 5.413744e-02 2.837204e-02 9.298240e-02 8.083232e-02
#> Locus_404 -7.779929e-02 -6.325941e-02 -3.353134e-02 -4.233244e-02
#> Locus_406 -2.637509e-01 1.004732e-01 1.049863e-02 1.888318e-01
#> Locus_409 -3.663654e-02 -1.824197e-02 -2.257098e-02 1.697614e-03
#> Locus_420 4.975221e-02 -1.977277e-02 2.143833e-02 -5.543596e-02
#> Locus_422 2.019295e-02 -7.925689e-03 2.355140e-02 -1.002941e-01
#> Locus_427 -4.197446e-02 -9.823366e-02 -2.108331e-02 -1.713880e-02
#> Locus_430 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_436 -3.172100e-03 5.439989e-02 2.159151e-02 -5.571553e-02
#> Locus_438 2.991668e-02 -5.177229e-02 -2.694501e-02 -6.160748e-02
#> Locus_439 -5.647847e-02 -8.088669e-02 6.358539e-02 3.400125e-02
#> Locus_441 -3.874442e-02 5.207570e-02 1.270215e-02 -1.846467e-01
#> Locus_445 -3.816226e-03 3.235327e-02 1.100763e-02 -4.333595e-02
#> Locus_448 4.203212e-02 4.196900e-02 4.291415e-03 -7.450235e-02
#> Locus_449 -2.621346e-02 -2.508331e-02 -1.479554e-02 -1.895013e-02
#> Locus_456 2.912308e-02 4.999338e-02 -3.318075e-02 -7.626164e-02
#> Locus_457 3.284994e-02 9.012221e-02 -1.839927e-02 -7.517258e-02
#> Locus_458 5.069119e-02 -1.947315e-02 1.972892e-03 -3.991466e-02
#> Locus_460 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_461 -3.813843e-02 9.974663e-03 9.300003e-03 -4.412196e-02
#> Locus_462 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_464 -5.923271e-02 -3.406833e-02 2.774252e-02 3.257056e-02
#> Locus_469 2.607776e-01 -1.828524e-01 2.790139e-02 -1.874152e-01
#> Locus_475 -1.867731e-02 3.250631e-02 -3.572980e-02 5.959791e-02
#> Locus_480 3.629349e-02 -4.114091e-02 -5.134661e-02 7.658834e-02
#> Locus_482 -2.102260e-02 -2.983385e-02 1.047556e-02 1.368527e-02
#> Locus_483 3.717849e-02 8.061383e-03 -2.980118e-02 3.794501e-02
#> Locus_491 -4.826132e-02 -1.790128e-02 -1.635443e-02 1.725986e-02
#> Locus_498 6.884952e-02 1.951092e-02 1.232133e-02 -1.759555e-01
#> Locus_500 1.152614e-02 -1.276289e-02 -8.423847e-03 -2.460324e-02
#> Locus_503 -4.396844e-02 3.141823e-02 -9.189391e-03 -1.126312e-01
#> Locus_505 -8.065897e-02 -3.150713e-02 4.228665e-02 -2.219876e-02
#> Locus_508 -1.087786e-01 -6.717974e-02 -2.709853e-02 -5.648897e-02
#> Locus_512 9.227459e-02 1.618075e-02 1.725376e-02 1.522059e-02
#> Locus_516 9.789693e-03 3.756064e-03 -1.503905e-02 1.074625e-02
#> Locus_517 -5.765909e-02 8.816409e-02 -4.740610e-03 9.877513e-02
#> Locus_518 7.314310e-02 -1.763285e-03 -4.036903e-03 -1.637218e-01
#> Locus_528 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_529 2.380445e-02 -4.167658e-02 3.445201e-02 -2.360839e-02
#> Locus_531 2.983225e-02 -5.439772e-03 8.474252e-03 4.468530e-02
#> Locus_532 -8.265122e-03 -4.149817e-04 4.573835e-02 5.153290e-03
#> Locus_533 -3.607932e-02 -7.922324e-03 -1.799902e-02 3.445015e-02
#> Locus_535 -2.898366e-02 1.613452e-02 3.257280e-02 1.362084e-03
#> Locus_546 -4.258946e-02 -2.327062e-02 4.525434e-02 -7.512452e-02
#> Locus_557 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_558 5.071457e-02 -4.042377e-03 4.249373e-02 -7.813934e-02
#> Locus_561 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_562 -5.527357e-02 2.646250e-02 2.742904e-03 -8.156154e-02
#> Locus_564 4.511646e-02 3.858017e-02 1.333677e-02 -1.019052e-01
#> Locus_566 -8.475694e-03 7.558315e-02 -1.045899e-02 9.088403e-03
#> Locus_572 -4.146199e-02 -1.219691e-03 -2.246129e-02 -9.097433e-02
#> Locus_573 -1.713513e-02 1.648784e-02 -6.557099e-03 -4.111417e-02
#> Locus_579 5.697572e-02 -2.302591e-02 -2.044612e-02 -1.313956e-01
#> Locus_581 -4.907211e-02 3.932495e-02 -2.422196e-02 -1.470868e-01
#> Locus_582 -1.713513e-02 1.648784e-02 -6.557099e-03 -4.111417e-02
#> Locus_583 -4.258069e-02 -1.477717e-03 2.328266e-02 8.384796e-02
#> Locus_586 -1.154406e-02 1.983788e-02 2.480951e-02 -8.216323e-02
#> Locus_589 -2.526766e-02 -2.351630e-03 1.023005e-02 -1.670206e-01
#> Locus_596 3.111098e-02 -4.373228e-02 -8.328318e-03 -8.192776e-02
#> Locus_597 5.657683e-02 -2.503316e-04 -5.324717e-03 7.999684e-03
#> Locus_598 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_600 3.095709e-02 -8.452460e-03 2.345810e-02 -5.484319e-02
#> Locus_602 -7.321479e-02 2.131959e-02 3.447064e-02 -3.834267e-02
#> Locus_603 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_604 -2.036498e-03 -2.174311e-02 2.616212e-03 1.200183e-02
#> Locus_609 -8.652717e-02 -4.763447e-02 -3.515535e-03 4.349095e-02
#> Locus_612 -2.952346e-02 -4.147601e-02 -1.107169e-02 3.836143e-02
#> Locus_613 3.321284e-03 2.128118e-02 5.791259e-02 2.322249e-02
#> Locus_614 3.905874e-02 -8.290478e-03 -2.281442e-03 -8.816601e-02
#> Locus_622 -2.388685e-02 -9.449683e-03 -6.580949e-03 6.202762e-03
#> Locus_625 -5.843406e-03 -1.877551e-03 2.950962e-02 1.851554e-02
#> Locus_626 -9.652264e-02 -3.580257e-02 -3.270886e-02 3.451973e-02
#> Locus_633 9.195272e-03 -1.950470e-02 -9.260582e-03 -1.850212e-02
#> Locus_634 1.190222e-02 -2.083829e-02 1.722601e-02 -1.180420e-02
#> Locus_635 -2.575064e-02 4.462534e-02 3.371848e-02 -1.626076e-01
#> Locus_636 3.663490e-01 -1.029604e-03 -7.841065e-02 -7.514924e-02
#> Locus_637 2.357052e-02 -2.265098e-02 7.022123e-03 -2.217244e-02
#> Locus_639 3.533188e-02 -4.841931e-02 -1.706292e-02 -8.052797e-02
#> Locus_642 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_647 -3.793328e-02 8.354907e-03 8.578681e-03 2.333546e-02
#> Locus_652 3.758154e-02 -2.298485e-02 7.936607e-03 -4.871061e-02
#> Locus_665 1.478146e-02 -7.272960e-04 -1.626908e-03 1.084394e-02
#> Locus_671 -4.681939e-02 2.117113e-03 -3.290217e-02 8.439566e-02
#> Locus_679 -8.535426e-03 -2.813518e-02 -2.150077e-02 -6.064696e-03
#> Locus_681 1.482377e-01 -1.900886e-02 -1.626044e-02 -2.494871e-01
#> Locus_682 -4.681939e-02 2.117113e-03 -3.290217e-02 8.439566e-02
#> Locus_683 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_685 -1.713513e-02 1.648784e-02 -6.557099e-03 -4.111417e-02
#> Locus_687 1.411783e-02 2.758060e-02 -1.132788e-02 -9.013269e-03
#> Locus_691 2.371398e-02 7.521060e-02 -1.954479e-04 -3.445205e-02
#> Locus_699 5.286707e-03 -8.636031e-02 -1.073229e-01 4.466629e-02
#> Locus_701 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_707 5.224388e-02 -3.513936e-02 -2.662531e-02 3.953151e-02
#> Locus_710 1.016458e-02 -1.387789e-02 -6.327616e-02 2.598669e-02
#> Locus_711 -3.384878e-02 -2.427216e-03 3.144030e-02 1.725129e-02
#> Locus_712 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_717 -8.535426e-03 -2.813518e-02 -2.150077e-02 -6.064696e-03
#> Locus_718 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_724 1.551167e-01 -3.639708e-02 1.438336e-02 1.225175e-02
#> Locus_725 1.190222e-02 -2.083829e-02 1.722601e-02 -1.180420e-02
#> Locus_727 3.952569e-02 -6.099041e-03 -9.248692e-03 2.864069e-02
#> Locus_729 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_733 -1.269361e-01 -1.212201e-02 4.931124e-04 9.772101e-02
#> Locus_734 1.461047e-02 -1.615172e-02 6.215093e-04 -4.471434e-02
#> Locus_735 2.157378e-02 -1.830335e-02 5.450449e-02 -6.364024e-02
#> Locus_736 -4.681939e-02 2.117113e-03 -3.290217e-02 8.439566e-02
#> Locus_740 -2.545441e-01 -1.351769e-01 -9.708459e-02 3.967321e-01
#> Locus_742 -2.004206e-02 4.414715e-02 -2.058926e-02 -5.135429e-03
#> Locus_746 3.434432e-02 1.695421e-02 8.984135e-02 2.090702e-02
#> Locus_750 -1.906922e-02 4.987332e-03 4.650002e-03 -2.206098e-02
#> Locus_751 -4.777371e-02 -1.889937e-02 -1.316190e-02 1.240552e-02
#> Locus_753 4.380982e-02 5.212037e-02 -3.503675e-02 -8.483377e-02
#> Locus_758 1.166829e-02 -1.812691e-03 -1.020388e-02 -1.036824e-02
#> Locus_765 4.929913e-03 2.540098e-02 -8.967837e-03 -1.918130e-02
#> Locus_769 -8.068856e-03 1.739731e-02 4.689292e-02 -1.957649e-02
#> Locus_770 2.854383e-01 -1.480320e-01 -5.428711e-02 -9.058983e-02
#> Locus_771 -1.785102e-01 -6.740715e-02 -1.029230e-01 1.278727e-01
#> Locus_776 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_779 -2.500440e-02 1.132786e-01 -3.611607e-02 1.483001e-02
#> Locus_784 -3.116647e-02 1.202711e-02 -2.268419e-02 2.650415e-02
#> Locus_786 -1.500633e-01 -1.044178e-02 -1.204412e-02 8.020838e-02
#> Locus_787 -2.351709e-02 -2.327303e-02 -2.397621e-02 3.505662e-02
#> Locus_798 1.015291e-01 6.566070e-02 -3.752494e-02 -3.621308e-01
#> Locus_802 -2.499337e-02 -5.807060e-04 3.020836e-02 8.121310e-02
#> Locus_803 -5.232907e-03 -4.350456e-03 1.066891e-02 -5.291836e-02
#> Locus_807 -3.813843e-02 9.974663e-03 9.300003e-03 -4.412196e-02
#> Locus_808 4.024778e-03 5.664824e-03 -4.924152e-02 -1.082591e-02
#> Locus_809 -4.777371e-02 -1.889937e-02 -1.316190e-02 1.240552e-02
#> Locus_811 3.766275e-02 -4.167750e-02 -1.622618e-02 -1.012126e-01
#> Locus_816 -3.942022e-03 3.409632e-02 -3.859990e-04 1.250141e-02
#> Locus_821 7.305237e-03 -8.075860e-03 3.107546e-04 -1.871130e-02
#> Locus_825 -1.932302e-02 -7.942822e-02 -3.552141e-02 -1.892485e-02
#> Locus_831 -9.341200e-02 3.643716e-02 1.204291e-02 -1.220089e-01
#> Locus_835 -1.082603e-02 -1.265447e-02 2.674801e-02 1.004056e-01
#> Locus_838 -6.664823e-02 -4.025470e-03 -2.229588e-02 9.418802e-02
#> Locus_847 -1.545871e-01 6.460436e-02 -8.852043e-03 3.541495e-02
#> Locus_850 -4.962341e-03 6.913142e-02 -1.552681e-02 1.735069e-02
#> Locus_851 6.708757e-02 -1.019424e-02 -1.496666e-02 5.204602e-02
#> Locus_853 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_854 -4.032948e-02 -1.575356e-02 2.114332e-02 -1.109938e-02
#> Locus_859 1.230328e-01 2.157433e-02 2.300501e-02 2.100864e-02
#> Locus_861 -3.060956e-03 1.051053e-01 -4.542242e-02 1.380217e-02
#> Locus_864 1.385949e-01 -5.929187e-03 1.200752e-02 1.079358e-01
#> Locus_869 -3.056891e-02 3.896854e-03 -4.296862e-03 6.420123e-02
#> Locus_870 -5.333948e-02 3.796300e-02 -8.464196e-03 -1.042893e-01
#> Locus_875 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_876 -5.517861e-02 3.471194e-02 -1.745400e-02 -6.048705e-02
#> Locus_879 8.984866e-02 -5.801826e-02 -1.606436e-02 -2.198729e-01
#> Locus_881 4.884276e-02 -1.168274e-01 6.626591e-02 1.066078e-01
#> Locus_887 4.639219e-02 1.002925e-02 3.377363e-02 3.267073e-02
#> Locus_888 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_895 1.660895e-02 -3.028483e-02 -3.680711e-02 1.885712e-02
#> Locus_896 1.264827e-01 -1.708860e-02 -9.746224e-03 1.150549e-01
#> Locus_897 3.127144e-02 1.779643e-02 -6.848702e-03 -4.767933e-03
#> Locus_898 -1.533778e-02 1.733920e-02 -1.709539e-02 -8.069782e-02
#> Locus_899 4.353030e-02 -2.730861e-02 2.616432e-02 5.058358e-02
#> Locus_901 -1.863788e-02 -1.018590e-01 -4.149108e-02 -3.787529e-02
#> Locus_902 -8.968176e-02 -2.242322e-02 -1.813035e-02 1.056634e-01
#> Locus_903 -9.366896e-02 2.220944e-02 1.267913e-02 -1.153887e-01
#> Locus_905 2.894739e-02 -4.560104e-02 3.882750e-03 -9.661071e-02
#> Locus_908 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_909 1.022280e-01 1.887822e-02 1.410329e-02 -1.072220e-01
#> Locus_915 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_924 5.524169e-02 4.326816e-02 -3.904098e-02 -8.428157e-03
#> Locus_932 -2.903114e-02 1.200980e-02 4.267125e-02 -1.101245e-02
#> Locus_935 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_936 -8.079369e-02 -6.596219e-02 1.032648e-01 -1.625789e-02
#> Locus_938 2.380445e-02 -4.167658e-02 3.445201e-02 -2.360839e-02
#> Locus_941 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_944 -1.961670e-02 -1.115713e-02 -4.994190e-03 8.884175e-03
#> Locus_945 -2.534378e-01 -1.200871e-02 -5.027900e-02 2.080220e-01
#> Locus_946 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_949 -3.224902e-02 -5.508229e-02 2.223881e-02 2.746379e-02
#> Locus_950 1.421308e-01 -1.384140e-04 -2.078737e-02 -3.342477e-01
#> Locus_951 9.261161e-02 -1.639107e-02 -8.197897e-03 8.986742e-03
#> Locus_953 -3.584563e-02 -7.161882e-02 -1.227554e-02 -3.162103e-02
#> Locus_958 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_965 3.085076e-02 -1.661639e-02 -2.384314e-02 -9.793955e-02
#> Locus_966 7.190625e-02 -6.277082e-03 -1.059869e-02 5.647116e-02
#> Locus_967 9.463818e-03 -2.436383e-02 -2.890841e-02 1.751288e-02
#> Locus_968 3.159301e-03 8.658327e-03 5.746826e-02 -3.806863e-03
#> Locus_970 2.529136e-02 2.006873e-02 -1.100973e-02 -4.569436e-02
#> Locus_972 2.974863e-02 -2.360622e-02 4.434435e-03 -5.242226e-02
#> Locus_974 -3.620435e-02 2.147517e-02 -1.907097e-03 -6.317515e-02
#> Locus_979 1.650051e-02 -2.758056e-02 -8.949828e-03 -3.592173e-02
#> Locus_992 3.758154e-02 -2.298485e-02 7.936607e-03 -4.583987e-02
#> Locus_998 -1.168681e-02 -3.755102e-03 5.901924e-02 3.703107e-02
#> Locus_999 3.402059e-04 -9.464625e-03 6.798718e-02 1.346105e-02
#> Locus_1002 4.876294e-03 1.866602e-02 -2.311459e-02 5.078807e-03
#> Locus_1005 1.747847e-01 -7.836696e-02 -8.733177e-02 1.373483e-01
#> Locus_1006 -2.721579e-02 2.745945e-02 2.674954e-02 -2.185915e-02
#> Locus_1007 9.305009e-02 -1.907053e-03 5.915242e-02 1.065161e-01
#> Locus_1015 4.602218e-02 8.799005e-03 2.125152e-02 4.477433e-02
#> Locus_1016 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1017 -7.098016e-02 1.271701e-02 -5.186740e-03 2.444868e-02
#> Locus_1023 1.190222e-02 -2.083829e-02 1.722601e-02 -1.180420e-02
#> Locus_1024 -7.812317e-03 5.551381e-02 -2.805152e-02 1.999806e-02
#> Locus_1025 -1.344644e-02 1.384725e-02 3.440271e-02 -3.320913e-02
#> Locus_1027 3.075820e-02 5.393583e-03 5.751253e-03 5.788050e-03
#> Locus_1030 3.017137e-01 -8.310645e-02 -8.288813e-02 -2.148412e-01
#> Locus_1031 3.388964e-02 -1.306248e-02 -6.177132e-02 4.042920e-02
#> Locus_1033 7.728280e-02 1.224234e-02 1.139129e-02 -1.171118e-01
#> Locus_1034 -8.207482e-02 1.976631e-02 -4.779630e-02 4.329870e-02
#> Locus_1035 2.956292e-02 -1.454592e-03 -3.253816e-03 2.222189e-02
#> Locus_1038 -2.036498e-03 -2.174311e-02 2.616212e-03 1.200183e-02
#> Locus_1041 3.186755e-02 6.764495e-02 4.437976e-02 -9.958688e-02
#> Locus_1042 -1.954287e-02 1.310994e-02 -3.071165e-02 1.359081e-02
#> Locus_1044 -8.838681e-02 -5.405951e-02 3.757017e-02 -2.739710e-02
#> Locus_1046 3.075820e-02 5.393583e-03 5.751253e-03 4.716269e-03
#> Locus_1051 6.151640e-02 1.078717e-02 1.150251e-02 1.050432e-02
#> Locus_1054 6.586574e-03 -2.380039e-02 -2.525107e-02 -2.603610e-02
#> Locus_1055 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1065 2.368775e-02 -2.829599e-03 -3.356745e-02 4.481919e-02
#> Locus_1071 2.333659e-02 -3.625381e-03 -2.040777e-02 -2.073649e-02
#> Locus_1072 5.171494e-02 1.227966e-01 1.951351e-04 -9.195391e-02
#> Locus_1073 -9.170501e-02 -2.136556e-02 -2.147791e-02 9.930574e-03
#> Locus_1074 6.054264e-02 -7.251004e-03 -9.635271e-03 4.038770e-02
#> Locus_1075 2.516732e-02 -2.274336e-02 2.302542e-02 -9.702907e-02
#> Locus_1076 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1080 2.086657e-02 -4.135969e-03 -1.518829e-02 -2.119726e-02
#> Locus_1082 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1093 3.955278e-02 -5.310634e-02 -2.579752e-02 -9.371164e-02
#> Locus_1095 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1098 -8.068856e-03 1.739731e-02 4.689292e-02 -1.957649e-02
#> Locus_1099 -9.432033e-03 1.683788e-03 1.964340e-03 2.269822e-02
#> Locus_1105 7.971078e-02 5.671866e-02 6.427761e-02 -7.041109e-02
#> Locus_1106 1.984080e-02 4.993843e-02 -2.995088e-04 -1.708485e-02
#> Locus_1107 2.444826e-02 7.861242e-03 -2.902951e-03 -4.345167e-02
#> Locus_1108 -3.427026e-02 3.297567e-02 -1.311420e-02 -8.222833e-02
#> Locus_1109 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1113 6.816873e-02 1.305439e-02 -3.510923e-03 -1.511117e-01
#> Locus_1115 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1118 -1.886407e-02 3.367576e-03 3.928679e-03 4.093308e-02
#> Locus_1122 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1127 -1.713513e-02 1.648784e-02 -6.557099e-03 -4.111417e-02
#> Locus_1138 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1141 -6.140834e-02 5.536031e-02 3.842872e-02 -1.201912e-01
#> Locus_1143 4.009366e-02 -4.086612e-02 -6.433748e-02 5.633592e-02
#> Locus_1144 4.408632e-02 -8.609467e-02 -3.673157e-02 -7.555293e-02
#> Locus_1155 8.746589e-03 -2.751466e-03 4.550924e-03 -2.417850e-03
#> Locus_1158 -1.794426e-02 1.089849e-01 -5.016139e-02 5.532944e-02
#> Locus_1159 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1161 4.716202e-02 -9.053371e-04 -2.976908e-03 3.814039e-02
#> Locus_1162 2.351372e-02 -4.760253e-02 -7.374101e-02 4.323764e-02
#> Locus_1166 3.579841e-02 -1.879259e-03 -2.013487e-03 2.259095e-02
#> Locus_1172 1.697980e-02 -1.001265e-01 4.092352e-03 -1.752784e-01
#> Locus_1179 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1181 3.075820e-02 5.393583e-03 5.751253e-03 4.716269e-03
#> Locus_1186 3.164295e-02 -4.156135e-02 -7.855309e-03 -3.717253e-02
#> Locus_1191 -2.921703e-03 -9.387754e-04 1.475481e-02 7.950394e-03
#> Locus_1197 2.441384e-02 -1.261272e-02 1.481680e-02 -9.889426e-02
#> Locus_1199 6.013977e-02 -3.694767e-03 1.460499e-02 -1.069549e-01
#> Locus_1201 -7.642207e-02 -6.291187e-02 -2.730666e-02 -3.291521e-02
#> Locus_1204 1.839054e-02 -3.900940e-02 -1.852116e-02 -2.842081e-02
#> Locus_1206 -3.116647e-02 1.202711e-02 -2.268419e-02 2.650415e-02
#> Locus_1211 -4.115573e-02 5.404310e-02 -8.584977e-03 -9.424939e-02
#> Locus_1212 3.493791e-02 7.073135e-03 -1.170772e-02 -5.452918e-02
#> Locus_1217 -3.890402e-03 2.077715e-02 6.846408e-02 -9.236645e-02
#> Locus_1223 4.203212e-02 4.196900e-02 4.291415e-03 -7.450235e-02
#> Locus_1224 4.158217e-02 1.309408e-02 2.644896e-02 1.765735e-02
#> Locus_1227 -1.741569e-02 9.771518e-02 -1.503455e-02 -2.346896e-02
#> Locus_1231 -3.427026e-02 3.297567e-02 -1.311420e-02 -8.222833e-02
#> Locus_1235 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1236 -2.442430e-02 -4.897100e-02 -5.865387e-02 -3.218408e-02
#> Locus_1239 -1.231978e-02 -1.856954e-02 5.025455e-02 6.196091e-02
#> Locus_1247 -1.254004e-01 -7.568124e-02 -6.721437e-02 -1.943912e-02
#> Locus_1248 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1249 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1250 3.685593e-02 3.993816e-02 -1.151770e-02 -1.002949e-01
#> Locus_1254 -3.620435e-02 2.147517e-02 -1.907097e-03 -6.317515e-02
#> Locus_1255 1.182445e-01 -2.318593e-02 8.082552e-02 2.834424e-02
#> Locus_1256 2.204234e-02 3.322121e-02 1.461679e-02 -5.437520e-02
#> Locus_1258 -6.198445e-03 -7.338961e-03 5.580049e-03 -1.486342e-01
#> Locus_1262 3.378973e-02 -1.386756e-02 -1.385036e-02 4.419395e-02
#> Locus_1263 -9.081796e-02 7.315905e-02 5.809586e-02 1.048165e-01
#> Locus_1265 3.095709e-02 -8.452460e-03 2.345810e-02 -5.340782e-02
#> Locus_1266 -6.421634e-02 -2.520325e-02 1.456238e-02 -4.896616e-03
#> Locus_1270 -2.402866e-02 -1.915298e-02 8.213420e-03 -8.148860e-03
#> Locus_1278 -2.070744e-01 -1.777080e-02 -3.702566e-02 2.110177e-01
#> Locus_1279 -8.061544e-02 -1.615702e-02 -2.764864e-02 7.742648e-02
#> Locus_1284 -5.884595e-02 1.192185e-02 6.241260e-03 7.972041e-02
#> Locus_1286 -2.544266e-02 3.902885e-02 -4.485968e-02 -1.158264e-01
#> Locus_1287 -1.584883e-01 -1.184804e-01 8.390380e-02 -2.238179e-01
#> Locus_1290 1.190222e-02 -2.083829e-02 1.722601e-02 -1.180420e-02
#> Locus_1295 1.513815e-02 -7.454495e-03 3.812926e-03 -1.499965e-02
#> Locus_1301 1.710346e-02 2.975917e-02 -4.453362e-02 -1.898693e-02
#> Locus_1302 -1.890823e-02 -3.315917e-02 -1.448479e-02 -4.391779e-02
#> Locus_1305 2.239226e-02 5.328365e-02 6.853854e-03 -2.125750e-03
#> Locus_1313 -4.533672e-03 4.148683e-02 -1.007299e-02 -6.027751e-03
#> Locus_1315 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1316 -9.952269e-02 4.096501e-02 -9.261377e-03 1.015854e-02
#> Locus_1318 -2.165064e-01 -1.608701e-02 -3.506132e-02 2.337159e-01
#> Locus_1319 -7.240870e-02 4.295033e-02 -3.814195e-03 -1.263503e-01
#> Locus_1338 -2.923690e-02 -8.813855e-03 1.001866e-02 1.002310e-01
#> Locus_1343 -2.825504e-02 2.312663e-02 5.190120e-02 -6.492463e-02
#> Locus_1345 5.641292e-02 -4.382360e-02 -1.764852e-04 -8.915441e-02
#> Locus_1347 1.951216e-02 -1.438222e-02 7.719156e-03 -6.815843e-02
#> Locus_1353 1.071932e-02 6.885277e-02 -3.715359e-03 -3.097730e-02
#> Locus_1354 4.621910e-02 1.842976e-02 2.472946e-02 3.646417e-02
#> Locus_1355 -1.235374e-02 7.450124e-04 1.671915e-02 3.064862e-02
#> Locus_1360 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1363 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1364 7.839560e-03 -3.479128e-02 9.187878e-03 -3.134418e-02
#> Locus_1366 1.371889e-02 8.076486e-02 -1.864732e-02 -4.028691e-02
#> Locus_1370 -4.228007e-02 -1.319811e-01 -3.609876e-02 -1.066923e-01
#> Locus_1374 -1.423065e-01 2.494842e-02 9.609715e-04 1.610337e-01
#> Locus_1376 -1.667116e-02 3.352493e-02 -1.283729e-02 -6.577582e-02
#> Locus_1377 -7.191727e-03 -3.419536e-02 -9.347094e-03 -8.746000e-03
#> Locus_1379 2.646447e-02 9.488218e-02 -6.341768e-02 -1.114094e-02
#> Locus_1380 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1382 2.338775e-02 3.998743e-02 2.497244e-02 -2.746850e-02
#> Locus_1384 -7.862680e-02 4.225332e-02 5.329504e-02 7.132755e-02
#> Locus_1392 1.308745e-01 1.588331e-02 -8.260915e-03 6.495472e-02
#> Locus_1396 -1.359080e-01 -1.040084e-01 4.936640e-02 -4.776016e-02
#> Locus_1399 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1404 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1408 1.126139e-01 -3.137849e-03 3.875562e-02 -2.222431e-01
#> Locus_1414 3.120438e-02 -1.576502e-02 2.026470e-02 -4.232893e-02
#> Locus_1416 -1.755009e-02 -4.630310e-02 -6.238339e-02 5.554559e-03
#> Locus_1420 -2.570913e-03 -3.577436e-02 3.623163e-03 1.913388e-02
#> Locus_1423 -7.185974e-02 4.029429e-02 1.048034e-02 -1.175163e-01
#> Locus_1424 -6.134545e-02 -1.440155e-02 -2.456523e-02 1.622315e-02
#> Locus_1425 5.099771e-02 -4.717658e-02 -1.005858e-02 -7.023393e-02
#> Locus_1426 6.770672e-02 -3.814408e-02 -1.045271e-02 1.733104e-01
#> Locus_1427 -2.388685e-02 -9.449683e-03 -6.580949e-03 6.202762e-03
#> Locus_1430 -9.695577e-02 1.664560e-02 -3.135534e-02 9.045785e-02
#> Locus_1433 -9.008516e-02 -1.354553e-03 6.256424e-02 -6.705513e-02
#> Locus_1439 -5.655957e-02 1.094514e-01 -5.832203e-02 9.522703e-02
#> Locus_1447 1.433041e-01 2.249339e-02 6.750291e-02 -6.876949e-02
#> Locus_1452 7.470217e-03 2.255491e-02 1.116853e-02 3.076929e-02
#> Locus_1455 9.878584e-02 -8.893000e-02 8.849545e-02 9.690087e-02
#> Locus_1457 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1459 -5.527357e-02 2.646250e-02 2.742904e-03 -7.788695e-02
#> Locus_1460 1.299635e-01 5.420239e-02 -3.846906e-02 -2.462281e-01
#> Locus_1461 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1463 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1468 6.227216e-02 9.764428e-02 -4.442706e-02 -1.720487e-01
#> Locus_1470 1.020251e-03 -4.249436e-02 -2.247224e-02 -2.530458e-01
#> Locus_1477 4.390950e-03 -2.339962e-02 -4.723950e-03 2.659769e-01
#> Locus_1479 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1482 9.195272e-03 -1.950470e-02 -9.260582e-03 -9.918692e-03
#> Locus_1485 -4.962341e-03 6.913142e-02 -1.552681e-02 1.735069e-02
#> Locus_1486 4.541446e-02 -2.236349e-02 1.143878e-02 -4.212821e-02
#> Locus_1487 6.854627e-02 5.422380e-02 4.673068e-03 -1.294585e-01
#> Locus_1488 -3.678735e-02 1.041898e-02 1.599617e-02 -1.919234e-03
#> Locus_1489 1.883137e-02 -2.083875e-02 -8.113092e-03 -4.331455e-02
#> Locus_1490 9.141802e-02 -4.255400e-02 1.932448e-02 -2.182502e-01
#> Locus_1494 4.700649e-02 2.715133e-02 3.765434e-03 -8.711253e-02
#> Locus_1496 -1.326826e-02 8.773497e-02 6.138814e-02 -9.641868e-02
#> Locus_1500 -3.060956e-03 1.051053e-01 -4.542242e-02 1.395131e-02
#> Locus_1502 -4.842166e-02 1.238399e-03 9.368037e-03 -3.871441e-03
#> Locus_1506 3.300102e-02 -5.516112e-02 -1.789966e-02 -5.725999e-02
#> Locus_1508 -1.755009e-02 -4.630310e-02 -6.238339e-02 5.554559e-03
#> Locus_1509 -4.785699e-03 6.590439e-03 7.985542e-02 8.532770e-03
#> Locus_1514 2.974863e-02 -2.360622e-02 4.434435e-03 -5.827862e-02
#> Locus_1515 3.947158e-02 -3.441369e-02 -1.634730e-03 -3.991800e-02
#> Locus_1517 2.842308e-02 8.249750e-03 2.097461e-02 2.832181e-02
#> Locus_1520 -2.193894e-02 -5.476297e-02 2.622161e-03 -4.810362e-02
#> Locus_1523 3.143765e-02 1.915705e-02 2.635636e-02 2.562023e-02
#> Locus_1524 -2.605223e-02 3.471390e-02 -6.180908e-02 -3.121571e-01
#> Locus_1529 -8.535426e-03 -2.813518e-02 -2.150077e-02 -6.064696e-03
#> Locus_1534 -5.959881e-03 2.399760e-02 -3.196803e-02 3.118158e-03
#> Locus_1536 -9.863712e-02 4.794510e-02 5.719068e-02 2.932544e-02
#> Locus_1540 7.480635e-02 1.965942e-02 6.186783e-02 6.915853e-02
#> Locus_1545 -1.979314e-01 -6.554662e-02 5.811297e-02 -1.444559e-01
#> Locus_1549 2.956292e-02 -1.454592e-03 -3.253816e-03 2.168788e-02
#> Locus_1551 -2.040637e-03 7.007019e-02 -3.028162e-02 9.102024e-03
#> Locus_1554 -5.834265e-02 -1.120618e-01 2.233569e-02 -1.711889e-02
#> Locus_1555 -7.812317e-03 5.551381e-02 -2.805152e-02 1.999806e-02
#> Locus_1561 2.291246e-02 -3.824718e-02 1.533581e-02 -7.072015e-02
#> Locus_1566 -2.073099e-02 -6.098455e-04 -1.123065e-02 -4.574601e-02
#> Locus_1572 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1573 -2.715016e-02 7.115436e-03 1.331051e-02 3.470202e-02
#> Locus_1576 1.215251e-02 -4.157642e-02 -3.739603e-02 3.044255e-02
#> Locus_1586 -6.615118e-02 -1.745663e-03 1.006630e-02 7.971731e-02
#> Locus_1587 2.072141e-02 -3.226759e-02 -1.768443e-02 -4.310536e-02
#> Locus_1590 -2.340970e-02 1.058556e-03 -1.645108e-02 4.219783e-02
#> Locus_1591 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1596 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1597 1.794285e-01 -4.799968e-02 -1.218128e-01 -2.667168e-02
#> Locus_1598 3.766275e-02 -4.167750e-02 -1.622618e-02 -1.012126e-01
#> Locus_1601 1.030986e-01 2.388125e-02 3.795146e-02 2.816167e-02
#> Locus_1602 -2.921703e-03 -9.387754e-04 1.475481e-02 7.950394e-03
#> Locus_1603 1.839054e-02 -3.900940e-02 -1.852116e-02 -2.842081e-02
#> Locus_1604 -1.190146e-02 4.155510e-02 -2.496000e-02 -1.366566e-02
#> Locus_1611 1.865331e-02 2.295840e-02 3.144173e-02 2.561049e-02
#> Locus_1612 -1.902174e-02 9.112053e-03 -5.448450e-03 -9.686441e-03
#> Locus_1614 -3.511454e-02 1.587835e-03 -2.467662e-02 6.559088e-02
#> Locus_1618 2.613661e-02 -2.891461e-02 -7.802338e-03 -7.660931e-02
#> Locus_1619 9.365358e-02 -8.886994e-02 -3.123568e-02 -2.581858e-01
#> Locus_1620 -5.934868e-02 -6.105386e-02 4.693204e-02 -6.707715e-03
#> Locus_1621 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1624 3.970125e-02 3.522719e-02 3.454679e-03 -6.840122e-02
#> Locus_1626 -2.921703e-03 -9.387754e-04 1.475481e-02 1.056514e-02
#> Locus_1632 -2.261760e-02 -7.985628e-03 -1.012200e-02 -1.431828e-02
#> Locus_1633 9.363617e-02 2.909381e-02 4.891825e-02 -3.339541e-02
#> Locus_1635 4.285356e-02 3.177506e-02 7.675073e-03 3.827359e-02
#> Locus_1636 -2.517180e-02 7.595635e-02 -3.628631e-02 8.727558e-03
#> Locus_1641 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1644 -1.346611e-01 -2.582791e-02 -2.340886e-02 -2.253056e-03
#> Locus_1645 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1646 -8.209355e-02 2.560688e-03 -7.598315e-04 1.099604e-01
#> Locus_1647 2.189797e-02 3.019222e-02 -1.943270e-02 -6.166192e-02
#> Locus_1648 6.327935e-02 7.297253e-02 1.867415e-03 -1.510034e-01
#> Locus_1649 3.833369e-03 -3.440984e-03 6.411892e-02 -3.138068e-02
#> Locus_1652 9.605255e-02 -1.064034e-01 -3.319066e-02 -1.396930e-01
#> Locus_1656 1.166829e-02 -1.812691e-03 -1.020388e-02 -1.036824e-02
#> Locus_1658 -2.696160e-02 6.095187e-03 -2.126613e-02 -6.930273e-02
#> Locus_1660 -1.284638e-02 1.373241e-01 -1.629881e-02 4.496825e-02
#> Locus_1661 1.774978e-02 1.175339e-02 3.581695e-02 -1.522129e-02
#> Locus_1664 6.476945e-02 -1.966398e-02 -2.023182e-02 6.182574e-02
#> Locus_1665 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1670 2.666429e-02 -2.021739e-02 -4.610921e-03 -3.960290e-02
#> Locus_1681 -2.996301e-02 -2.110463e-02 -9.801419e-03 1.919029e-02
#> Locus_1682 -1.080575e-02 6.725387e-02 1.398281e-02 3.556795e-02
#> Locus_1687 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1690 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1691 -2.490361e-02 -4.196330e-02 2.191823e-02 -6.044780e-03
#> Locus_1692 2.474423e-02 -5.371745e-03 -7.621784e-03 1.779675e-02
#> Locus_1693 -7.622167e-02 1.993151e-03 3.918439e-02 -3.280490e-02
#> Locus_1694 -9.102270e-02 -1.247707e-02 -3.206937e-02 1.097085e-01
#> Locus_1697 1.082789e-01 -6.631038e-02 5.182635e-04 -1.863626e-01
#> Locus_1699 6.795990e-02 -8.421967e-02 -1.647975e-02 -1.116659e-01
#> Locus_1702 3.705386e-02 -3.168208e-02 4.745190e-03 -7.698992e-02
#> Locus_1703 -1.429913e-02 -1.637273e-02 -1.247098e-02 4.188725e-02
#> Locus_1708 1.985237e-02 -4.896281e-02 1.129921e-02 -2.264790e-02
#> Locus_1713 3.837717e-02 1.829212e-02 2.631356e-02 -8.206902e-02
#> Locus_1715 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1724 -1.981912e-01 7.093887e-03 2.741169e-02 2.138298e-01
#> Locus_1728 -8.103439e-02 -3.100499e-02 6.138197e-02 7.782880e-02
#> Locus_1730 2.990173e-02 -2.584691e-02 1.707527e-02 -1.142726e-01
#> Locus_1734 -7.041571e-02 -9.351459e-03 1.024874e-02 5.196751e-02
#> Locus_1737 -4.227376e-02 4.426132e-03 -1.252240e-02 8.300601e-02
#> Locus_1742 -8.090784e-02 1.335599e-03 1.818334e-02 1.343970e-01
#> Locus_1745 -7.240870e-02 4.295033e-02 -3.814195e-03 -1.263503e-01
#> Locus_1748 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1750 -2.824759e-02 -1.641996e-02 1.588805e-03 -6.442810e-02
#> Locus_1751 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1755 1.839054e-02 -3.900940e-02 -1.852116e-02 -3.700424e-02
#> Locus_1756 4.234334e-02 -4.822490e-03 -7.344876e-03 3.424927e-02
#> Locus_1757 -4.032948e-02 -1.575356e-02 2.114332e-02 -1.109938e-02
#> Locus_1774 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1776 9.307793e-02 -2.556922e-02 2.207901e-02 -1.273516e-02
#> Locus_1777 2.974863e-02 -2.360622e-02 4.434435e-03 -6.557035e-02
#> Locus_1778 -2.040637e-03 7.007019e-02 -3.028162e-02 9.251160e-03
#> Locus_1783 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1784 -8.991999e-02 -8.882839e-02 1.452053e-02 -6.976502e-02
#> Locus_1786 2.705735e-02 -3.417220e-02 1.345409e-02 -8.823646e-02
#> Locus_1787 5.798600e-02 6.795158e-03 1.772080e-02 5.000969e-02
#> Locus_1797 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1799 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1801 -1.742327e-01 -2.051314e-02 -2.253892e-02 1.507099e-01
#> Locus_1804 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1805 6.201454e-03 1.286245e-02 -2.696487e-02 -6.185066e-02
#> Locus_1807 1.245339e-01 -6.289468e-02 3.969527e-03 -2.854601e-01
#> Locus_1808 1.536014e-01 -2.176694e-02 7.898860e-03 4.433944e-02
#> Locus_1811 -2.624731e-02 -3.590311e-02 9.764553e-03 -2.845791e-03
#> Locus_1813 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1816 2.842308e-02 8.249750e-03 2.097461e-02 2.832181e-02
#> Locus_1822 -1.460432e-02 7.599252e-02 -4.096223e-02 3.559425e-02
#> Locus_1823 -3.474889e-02 -5.321849e-02 -3.629631e-02 -2.397945e-02
#> Locus_1826 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1828 3.061229e-02 3.210331e-02 8.981115e-03 1.434553e-02
#> Locus_1833 -2.202851e-01 -2.530174e-02 -5.229310e-03 2.397869e-01
#> Locus_1839 1.957939e-02 7.512128e-03 -3.007809e-02 2.149250e-02
#> Locus_1840 -5.917387e-02 -8.987626e-03 8.804030e-03 -8.827311e-02
#> Locus_1841 1.346179e-01 1.135826e-02 9.908882e-03 4.946985e-02
#> Locus_1842 3.547274e-02 -4.348927e-02 2.424813e-02 -3.397664e-02
#> Locus_1843 4.180244e-02 -2.767188e-02 -7.979945e-04 -5.173180e-02
#> Locus_1844 4.352361e-03 -1.608074e-02 1.153663e-02 -1.705228e-02
#> Locus_1846 -2.437447e-02 -8.451602e-03 -9.773481e-03 1.105710e-02
#> Locus_1848 -9.785428e-03 3.221877e-02 2.912362e-02 3.101694e-02
#> Locus_1849 -3.582113e-02 3.059199e-02 5.239121e-03 -9.778860e-02
#> Locus_1852 9.956048e-02 1.419050e-01 1.329494e-02 -2.436005e-01
#> Locus_1853 1.927078e-02 -1.897554e-04 2.437902e-02 -2.963614e-02
#> Locus_1856 2.666429e-02 -2.021739e-02 -4.610921e-03 -3.960290e-02
#> Locus_1861 3.396953e-02 -2.829325e-02 -4.300166e-03 -5.687883e-02
#> Locus_1863 -9.432033e-03 1.683788e-03 1.964340e-03 1.823486e-02
#> Locus_1868 -8.591214e-02 -8.924702e-03 -3.861895e-03 -2.436726e-02
#> Locus_1870 -2.921703e-03 -9.387754e-04 1.475481e-02 7.950394e-03
#> Locus_1871 -7.562962e-03 -2.087837e-02 1.661680e-02 -3.131925e-02
#> Locus_1872 -7.834262e-02 -1.500251e-02 -1.745876e-02 8.119502e-02
#> Locus_1873 1.814745e-01 -9.802106e-02 6.984459e-03 -3.521915e-01
#> Locus_1881 1.083063e-01 3.006104e-02 -4.257941e-02 -6.335408e-02
#> Locus_1882 -1.771813e-02 5.431648e-03 1.134617e-02 1.646716e-02
#> Locus_1887 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1894 -3.544583e-02 1.079152e-02 -5.442425e-02 -1.777815e-02
#> Locus_1896 1.839054e-02 -3.900940e-02 -1.852116e-02 -2.842081e-02
#> Locus_1908 -7.705772e-03 9.588672e-02 1.151852e-02 -6.174328e-02
#> Locus_1911 -3.772813e-02 6.735151e-03 7.857359e-03 8.186617e-02
#> Locus_1913 3.163788e-02 -1.995929e-03 3.929035e-02 -7.968708e-02
#> Locus_1919 -1.902174e-02 9.112053e-03 -5.448450e-03 -9.686441e-03
#> Locus_1921 4.552930e-02 1.245695e-02 1.398348e-02 -5.494886e-02
#> Locus_1932 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1933 -2.504898e-02 1.588212e-02 -6.927388e-03 -3.448712e-03
#> Locus_1935 -1.713513e-02 1.648784e-02 -6.557099e-03 -4.111417e-02
#> Locus_1939 1.581894e-02 -9.979645e-04 1.964517e-02 -3.840817e-02
#> Locus_1941 1.649972e-02 5.458566e-03 3.547742e-02 -6.325206e-02
#> Locus_1942 1.152614e-02 -1.276289e-02 -8.423847e-03 -2.460324e-02
#> Locus_1945 -9.405522e-03 -4.570391e-02 4.381778e-02 -1.321713e-02
#> Locus_1948 -7.191727e-03 -3.419536e-02 -9.347094e-03 -8.746000e-03
#> Locus_1949 -1.193036e-01 -6.536143e-02 -5.639121e-02 1.607096e-01
#> Locus_1950 1.661456e-02 4.027900e-02 3.802213e-02 -7.463733e-02
#> Locus_1951 2.321491e-02 -2.985339e-02 6.952472e-03 -5.146071e-02
#> Locus_1952 -5.127076e-02 -4.597024e-02 1.679036e-02 1.777735e-02
#> Locus_1953 -1.992385e-02 1.404493e-02 7.604249e-02 2.209705e-02
#> Locus_1958 4.844987e-02 -2.373630e-02 -1.649598e-02 3.750711e-02
#> Locus_1961 2.301673e-02 -1.068625e-01 2.785576e-02 -1.691679e-01
#> Locus_1962 -1.931158e-01 -1.012690e-01 6.896787e-02 -1.572252e-01
#> Locus_1965 8.184297e-03 -2.084076e-02 -3.292116e-02 2.094670e-02
#> Locus_1967 -3.427026e-02 3.297567e-02 -1.311420e-02 -8.222833e-02
#> Locus_1968 1.649972e-02 5.458566e-03 3.547742e-02 -6.325206e-02
#> Locus_1971 4.537301e-02 1.621640e-02 4.290024e-02 2.931650e-02
#> Locus_1974 1.431026e-02 -2.057461e-02 -3.169323e-02 1.893670e-02
#> Locus_1977 -6.663777e-02 -1.553179e-02 -9.233219e-03 4.887525e-02
#> Locus_1982 3.306683e-02 2.595015e-02 -6.101566e-02 -3.378578e-02
#> Locus_1983 1.936257e-02 8.499421e-02 -2.349212e-02 2.157396e-02
#> Locus_1984 1.321235e-02 9.743614e-02 -2.392399e-02 -1.569839e-02
#> Locus_1988 -3.813843e-02 9.974663e-03 9.300003e-03 -4.044737e-02
#> Locus_1990 -8.791045e-03 -3.674500e-02 5.894149e-02 -3.609920e-02
#> Locus_1991 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1993 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2003 1.864101e-02 -3.256545e-02 -2.167463e-02 -9.631553e-02
#> Locus_2007 -3.427026e-02 3.297567e-02 -1.311420e-02 -8.222833e-02
#> Locus_2014 9.227459e-02 1.618075e-02 1.725376e-02 1.522059e-02
#> Locus_2016 -9.652264e-02 -3.580257e-02 -3.270886e-02 3.451973e-02
#> Locus_2017 4.464866e-02 -1.837352e-02 3.289685e-02 3.728014e-02
#> Locus_2019 -3.126967e-02 -1.784315e-02 6.461302e-02 9.635588e-03
#> Locus_2020 1.422495e-02 1.932675e-02 -1.329729e-02 2.666005e-02
#> Locus_2021 -1.394848e-02 2.009681e-02 -4.327106e-02 -1.533562e-03
#> Locus_2027 -4.032948e-02 -1.575356e-02 2.114332e-02 -1.109938e-02
#> Locus_2030 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2033 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2036 -3.942022e-03 3.409632e-02 -3.859990e-04 1.511615e-02
#> Locus_2041 -4.962341e-03 6.913142e-02 -1.552681e-02 1.735069e-02
#> Locus_2045 -2.194281e-01 -1.702578e-02 -2.030651e-02 2.416663e-01
#> Locus_2048 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2050 2.380445e-02 -4.167658e-02 3.445201e-02 -2.360839e-02
#> Locus_2053 -1.941697e-01 -6.400125e-03 1.162318e-01 1.783341e-01
#> Locus_2055 -1.023650e-01 1.645162e-02 -2.101045e-02 2.603363e-02
#> Locus_2060 1.792868e-02 3.340464e-02 -6.482859e-02 -1.878612e-02
#> Locus_2063 -1.906922e-02 4.987332e-03 4.650002e-03 -1.838639e-02
#> Locus_2064 8.828960e-02 1.397130e-02 4.949506e-02 -2.100309e-02
#> Locus_2068 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2069 -5.771908e-02 1.101229e-02 9.013649e-03 1.034915e-01
#> Locus_2080 -5.022788e-02 -2.793304e-02 -2.761263e-02 2.858463e-02
#> Locus_2085 2.444826e-02 7.861242e-03 -2.902951e-03 -4.345167e-02
#> Locus_2086 2.333659e-02 -3.625381e-03 -2.040777e-02 -2.073649e-02
#> Locus_2087 1.166829e-02 -1.812691e-03 -1.020388e-02 -1.036824e-02
#> Locus_2091 2.408462e-02 -6.247130e-03 -1.775967e-02 -5.399224e-02
#> Locus_2094 4.388033e-03 -7.195088e-02 8.381366e-03 -6.442021e-02
#> Locus_2102 -4.768278e-02 -4.241976e-02 8.861156e-03 -8.151618e-02
#> Locus_2106 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2108 -3.772813e-02 6.735151e-03 7.857359e-03 8.186617e-02
#> Locus_2109 -9.432033e-03 1.683788e-03 1.964340e-03 2.269822e-02
#> Locus_2113 1.448763e-02 -4.443259e-02 -5.261939e-02 7.908792e-03
#> Locus_2115 1.650051e-02 -2.758056e-02 -8.949828e-03 -3.721342e-02
#> Locus_2120 9.551065e-02 -1.944485e-03 6.007953e-03 8.036783e-02
#> Locus_2128 7.368553e-03 2.064854e-02 7.153018e-03 -1.783195e-02
#> Locus_2131 1.177327e-01 3.259557e-02 6.101123e-03 -1.961876e-01
#> Locus_2133 -9.147791e-02 4.793767e-02 8.358071e-04 -1.410621e-01
#> Locus_2135 9.195272e-03 -1.950470e-02 -9.260582e-03 -9.918692e-03
#> Locus_2136 1.091298e-01 5.502271e-02 -2.887717e-03 -2.798585e-01
#> Locus_2137 -5.430032e-02 1.423087e-02 2.662102e-02 6.940404e-02
#> Locus_2138 -2.490361e-02 -4.196330e-02 2.191823e-02 -6.044780e-03
#> Locus_2139 -1.906922e-02 4.987332e-03 4.650002e-03 -2.206098e-02
#> Locus_2140 3.243429e-02 6.241913e-03 1.324005e-02 -9.429859e-02
#> Locus_2145 -2.961442e-02 3.877841e-02 -6.959361e-03 -8.984680e-02
#> Locus_2148 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2154 -5.935122e-02 -6.641511e-03 1.569487e-02 -2.078582e-02
#> Locus_2157 1.975212e-02 -2.609634e-02 1.314333e-02 -8.669202e-02
#> Locus_2159 6.504923e-02 2.180067e-03 3.286579e-03 4.533754e-02
#> Locus_2162 5.263070e-02 -2.673879e-04 -1.730529e-02 5.301851e-03
#> Locus_2165 2.758581e-02 -5.851410e-02 -2.778175e-02 -3.833951e-02
#> Locus_2166 -6.791998e-03 2.047871e-02 -1.291071e-02 1.544705e-02
#> Locus_2168 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2179 -1.374549e-02 7.059947e-02 -3.850716e-02 3.279334e-02
#> Locus_2180 1.463207e-01 -8.759050e-03 -1.272185e-02 1.061936e-01
#> Locus_2181 7.791155e-02 -2.493740e-03 5.731045e-03 6.391531e-02
#> Locus_2183 -2.540074e-03 3.060054e-02 1.460924e-02 -1.214716e-01
#> Locus_2185 -2.379338e-02 1.049878e-02 -4.362387e-02 -6.324735e-02
#> Locus_2186 4.716202e-02 -9.053371e-04 -2.976908e-03 3.920842e-02
#> Locus_2188 1.525299e-02 2.736594e-02 6.357631e-03 -2.494955e-02
#> Locus_2191 2.255823e-02 1.929008e-02 6.668385e-03 -5.095259e-02
#> Locus_2203 5.621516e-02 -5.345357e-05 1.672379e-04 4.411802e-02
#> Locus_2208 -1.431910e-02 7.370560e-02 1.905063e-02 2.498919e-02
#> Locus_2211 4.689618e-02 1.917742e-01 -1.385017e-02 -1.791497e-01
#> Locus_2213 6.234111e-02 2.126978e-04 1.395164e-03 4.210802e-02
#> Locus_2215 -1.069975e-02 1.032469e-02 -6.092076e-03 -1.044008e-01
#> Locus_2216 3.897805e-02 2.818135e-02 7.862466e-03 -1.050704e-02
#> Locus_2219 -3.853832e-02 6.202442e-03 -5.037274e-02 -1.310905e-01
#> Locus_2223 3.039114e-02 1.991145e-02 1.017056e-02 -3.851383e-02
#> Locus_2228 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2229 -3.595863e-03 -1.709768e-02 -4.673547e-03 -4.631843e-03
#> Locus_2230 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2231 7.305237e-03 -8.075860e-03 3.107546e-04 -1.871130e-02
#> Locus_2234 1.513815e-02 -7.454495e-03 3.812926e-03 -1.356428e-02
#> Locus_2236 -3.118687e-02 -4.694061e-02 -7.062670e-03 -4.278645e-03
#> Locus_2238 3.830526e-02 1.840163e-03 -1.049006e-02 -7.415604e-02
#> Locus_2243 6.070809e-03 7.551916e-02 -1.633305e-02 -2.705515e-02
#> Locus_2246 1.526773e-02 1.377193e-02 2.648515e-02 -4.031297e-02
#> Locus_2250 6.665645e-02 9.873284e-02 -2.632963e-02 -9.521877e-02
#> Locus_2251 9.975533e-03 -2.875515e-03 4.915479e-02 -2.132800e-02
#> Locus_2254 1.296686e-02 -5.027219e-02 6.758334e-04 -4.200182e-02
#> Locus_2260 -5.184296e-04 3.523426e-05 -7.416483e-03 1.609350e-02
#> Locus_2261 3.533059e-02 -5.443947e-02 2.602817e-02 -4.821164e-02
#> Locus_2267 -7.367539e-02 -6.314340e-02 2.264763e-02 8.896283e-02
#> Locus_2268 7.846825e-02 6.740218e-02 -6.854205e-02 -1.486968e-01
#> Locus_2271 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2273 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2274 -5.186227e-02 -5.874658e-02 -7.317714e-02 2.712829e-02
#> Locus_2275 6.872694e-02 -2.809177e-02 -4.255643e-02 6.080039e-02
#> Locus_2277 1.445623e-02 -1.879437e-02 3.204832e-02 1.447491e-02
#> Locus_2282 9.320332e-02 5.811139e-02 7.697746e-02 -7.179747e-02
#> Locus_2287 6.741881e-02 -2.688645e-02 4.515759e-02 1.287726e-01
#> Locus_2294 6.201454e-03 1.286245e-02 -2.696487e-02 -6.185066e-02
#> Locus_2298 9.789693e-03 3.756064e-03 -1.503905e-02 1.074625e-02
#> Locus_2305 1.924038e-01 1.883573e-02 -2.868891e-02 -3.484830e-01
#> Locus_2308 -4.928164e-02 1.713381e-02 -3.149524e-02 2.196001e-02
#> Locus_2309 -2.340970e-02 1.058556e-03 -1.645108e-02 4.219783e-02
#> Locus_2312 1.088836e-01 -1.763785e-02 -1.002313e-02 1.030347e-01
#> Locus_2318 2.679428e-02 -4.848499e-02 -6.726411e-02 3.862039e-02
#> Locus_2319 3.338967e-02 -9.864355e-03 6.444146e-02 1.537504e-02
#> Locus_2321 8.821518e-02 -4.034628e-02 -1.051146e-01 4.319854e-02
#> Locus_2322 -4.349698e-02 7.707827e-03 -4.230850e-02 -1.196698e-01
#> Locus_2323 2.651270e-02 -3.699001e-02 1.784752e-02 -6.381026e-02
#> Locus_2325 1.573559e-02 -2.427928e-02 8.134493e-02 -4.318488e-02
#> Locus_2326 3.556821e-02 2.328750e-03 1.307592e-02 2.966605e-02
#> Locus_2327 -1.450024e-02 -2.257918e-06 4.478427e-02 -1.156250e-01
#> Locus_2333 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2336 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2338 -3.961632e-01 4.005715e-02 -6.826325e-02 2.843730e-01
#> Locus_2339 3.039114e-02 1.991145e-02 1.017056e-02 -3.851383e-02
#> Locus_2344 -5.792063e-02 3.905173e-02 -2.113394e-02 1.190254e-01
#> Locus_2345 -1.771813e-02 5.431648e-03 1.134617e-02 1.646716e-02
#> Locus_2348 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2349 1.839054e-02 -3.900940e-02 -1.852116e-02 -1.983738e-02
#> Locus_2354 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2359 1.221645e-02 -8.393270e-03 1.856774e-02 -5.613887e-03
#> Locus_2361 -6.177485e-02 -5.554773e-02 -2.091633e-02 1.091662e-01
#> Locus_2364 -3.430913e-02 -8.766720e-02 6.573601e-02 -1.926191e-02
#> Locus_2365 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2366 1.284837e-02 5.874607e-02 -1.921371e-02 -8.277287e-02
#> Locus_2368 -3.267323e-02 8.779491e-02 4.688819e-05 4.007234e-02
#> Locus_2371 -2.040637e-03 7.007019e-02 -3.028162e-02 9.102024e-03
#> Locus_2375 -7.151636e-02 -6.269418e-02 1.408065e-02 -1.537802e-02
#> Locus_2381 -7.191727e-03 -3.419536e-02 -9.347094e-03 -8.746000e-03
#> Locus_2388 -6.278759e-03 3.288156e-02 -2.551066e-02 1.218280e-02
#> Locus_2394 -3.596475e-02 -2.629220e-02 -7.295323e-02 -3.212579e-02
#> Locus_2400 1.402697e-01 -1.460695e-02 -1.336231e-02 9.219752e-02
#> Locus_2402 2.504097e-03 5.038047e-02 -3.932591e-02 -4.060052e-03
#> Locus_2406 3.276879e-02 -2.497426e-02 2.037719e-03 -3.300145e-02
#> Locus_2413 -3.380650e-02 -6.767814e-03 -7.809141e-03 2.929196e-02
#> Locus_2414 -3.942022e-03 3.409632e-02 -3.859990e-04 1.511615e-02
#> Locus_2415 -1.905463e-02 2.584908e-02 -2.799790e-02 4.533247e-02
#> Locus_2421 -3.620435e-02 2.147517e-02 -1.907097e-03 -5.950056e-02
#> Locus_2424 -4.962341e-03 6.913142e-02 -1.552681e-02 1.705242e-02
#> Locus_2425 2.332965e-01 -1.322728e-02 -1.896087e-02 1.642533e-01
#> Locus_2426 -5.333948e-02 3.796300e-02 -8.464196e-03 -1.042893e-01
#> Locus_2431 -1.841970e-01 -2.626932e-02 -4.227754e-02 1.502013e-01
#> Locus_2432 -2.040637e-03 7.007019e-02 -3.028162e-02 9.251160e-03
#> Locus_2433 -4.746828e-02 2.332847e-03 -7.957476e-03 9.426598e-02
#> Locus_2434 -3.427026e-02 3.297567e-02 -1.311420e-02 -8.222833e-02
#> Locus_2435 -9.432033e-03 1.683788e-03 1.964340e-03 1.823486e-02
#> Locus_2436 -1.874595e-02 -4.009369e-02 -9.051894e-02 1.074843e-01
#> Locus_2437 -3.772813e-02 6.735151e-03 7.857359e-03 8.186617e-02
#> Locus_2439 -4.287758e-03 4.260036e-03 3.023094e-03 -7.542455e-03
#> Locus_2448 -1.089289e-01 -1.035836e-01 -1.632526e-02 -2.168641e-01
#> Locus_2449 -1.743813e-02 3.753836e-02 -4.856028e-03 -1.132138e-01
#> Locus_2450 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2451 8.068328e-02 -3.470923e-02 -2.482890e-02 -1.453325e-01
#> Locus_2452 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2457 -7.827602e-02 3.142174e-02 -5.259773e-02 -1.753313e-01
#> Locus_2461 -2.113688e-02 2.213066e-03 -6.261202e-03 4.150301e-02
#> Locus_2463 1.977002e-02 -3.297793e-02 5.789847e-02 -3.339664e-02
#> Locus_2466 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2467 -1.066301e-01 4.528807e-03 -1.998330e-02 1.283871e-01
#> Locus_2473 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2475 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2477 -1.026213e-01 -1.055134e-01 -4.875114e-02 6.782761e-02
#> Locus_2480 1.879991e-01 -6.301433e-03 1.756281e-03 1.299953e-01
#> Locus_2494 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2497 6.403468e-02 8.267989e-03 -1.992977e-03 -1.004676e-01
#> Locus_2501 2.983091e-02 1.256635e-02 -4.760258e-02 -3.059032e-02
#> Locus_2502 2.305227e-02 -2.552578e-02 -1.684769e-02 -4.920649e-02
#> Locus_2507 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2510 4.639219e-02 1.002925e-02 3.377363e-02 4.153534e-02
#> Locus_2512 7.179697e-02 3.212930e-02 -4.781874e-02 -2.570104e-01
#> Locus_2514 -2.452527e-02 1.074074e-02 -3.017439e-02 5.977083e-02
#> Locus_2517 1.006993e-01 3.202909e-03 2.289802e-02 8.208800e-02
#> Locus_2518 6.284918e-03 2.695924e-02 -1.483005e-02 -1.416029e-02
#> Locus_2519 -2.066672e-02 -7.336804e-02 -4.767509e-02 -1.572586e-02
#> Locus_2522 2.716995e-02 -7.066365e-03 4.371115e-02 -5.211717e-02
#> Locus_2526 -6.113783e-02 7.793708e-03 -8.593725e-03 1.194757e-01
#> Locus_2529 -2.261760e-02 -7.985628e-03 -1.012200e-02 -1.380060e-02
#> Locus_2530 -8.904362e-03 1.032277e-01 -1.591281e-02 2.985210e-02
#> Locus_2532 6.977626e-02 -2.740683e-02 -2.345428e-02 6.054338e-02
#> Locus_2540 -3.171276e-02 6.213632e-03 -1.611583e-02 -6.387873e-02
#> Locus_2542 -5.843406e-03 -1.877551e-03 2.950962e-02 1.851554e-02
#> Locus_2546 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2550 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2555 -5.430032e-02 1.423087e-02 2.662102e-02 7.833076e-02
#> Locus_2559 -1.730660e-01 3.163130e-02 9.171677e-02 2.150980e-01
#> Locus_2561 -7.755230e-02 -4.069687e-02 -1.311612e-02 -6.363328e-02
#> Locus_2565 1.190222e-02 -2.083829e-02 1.722601e-02 -1.180420e-02
#> Locus_2566 -3.427026e-02 3.297567e-02 -1.311420e-02 -8.222833e-02
#> Locus_2567 -5.718851e-02 -4.028938e-02 -4.635996e-02 1.689971e-02
#> Locus_2572 -4.681939e-02 2.117113e-03 -3.290217e-02 8.439566e-02
#> Locus_2573 4.964521e-02 -8.363509e-02 -1.670223e-02 -1.632932e-01
#> Locus_2577 5.963300e-02 -1.754672e-03 -4.962497e-04 3.887850e-02
#> Locus_2579 -1.005553e-01 -1.195941e-01 9.586343e-02 -4.033914e-02
#> Locus_2581 -6.441789e-02 -9.206410e-03 9.429482e-03 -1.838827e-01
#> Locus_2583 1.161238e-04 -5.400094e-02 2.505994e-02 1.732580e-02
#> Locus_2588 1.622157e-01 1.399008e-02 3.440053e-02 9.152054e-02
#> Locus_2589 -5.985891e-02 1.283130e-02 9.700583e-03 1.654783e-01
#> Locus_2592 -3.823196e-02 -5.874587e-02 6.824911e-02 -2.087851e-02
#> Locus_2597 -6.791998e-03 2.047871e-02 -1.291071e-02 1.544705e-02
#> Locus_2598 4.669082e-02 -2.075265e-02 -3.304323e-02 4.676717e-02
#> Locus_2599 1.166829e-02 -1.812691e-03 -1.020388e-02 -1.036824e-02
#> Locus_2601 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2602 -1.020740e-01 -4.668493e-03 1.345912e-02 8.180956e-02
#> Locus_2611 3.247193e-02 2.310149e-02 6.209311e-02 2.674328e-02
#> Locus_2616 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2617 9.377496e-03 1.774501e-02 -1.874216e-03 2.588248e-02
#> Locus_2619 2.974863e-02 -2.360622e-02 4.434435e-03 -6.557035e-02
#> Locus_2620 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2629 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2632 -1.448618e-02 -1.062825e-02 6.772653e-02 -7.769582e-02
#> Locus_2634 2.815457e-02 2.312550e-02 -4.678024e-02 -8.379086e-02
#> Locus_2636 -1.864321e-03 -1.215954e-02 -8.173847e-03 2.415380e-02
#> Locus_2639 -3.817820e-02 -1.623095e-02 5.008129e-02 -1.343201e-02
#> Locus_2644 6.406944e-02 -2.825797e-02 -3.199436e-02 -9.106214e-02
#> Locus_2647 1.179741e-01 2.767954e-03 2.297756e-02 -2.803722e-01
#> Locus_2648 2.677913e-02 1.460305e-02 -2.066216e-03 -4.955280e-02
#> Locus_2652 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2655 2.668368e-02 -2.156559e-02 1.559910e-02 -9.602586e-04
#> Locus_2656 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2657 -1.177121e-01 -1.462017e-02 -2.509178e-02 -4.601111e-03
#> Locus_2660 3.533188e-02 -4.841931e-02 -1.706292e-02 -8.652800e-02
#> Locus_2662 -1.135985e-01 1.997064e-02 -5.036229e-03 2.224861e-01
#> Locus_2665 -1.555535e-01 -5.266714e-02 -3.983591e-02 -4.284163e-02
#> Locus_2668 5.068217e-02 5.950729e-02 -6.256261e-02 1.094884e-02
#> Locus_2669 -6.788671e-02 3.748029e-02 2.846307e-02 -1.691392e-01
#> Locus_2670 -7.644312e-02 5.164899e-02 1.963226e-02 -1.324639e-01
#> Locus_2671 1.166829e-02 -1.812691e-03 -1.020388e-02 -1.036824e-02
#> Locus_2675 1.917886e-02 -1.165079e-01 -9.402331e-03 2.580160e-02
#> Locus_2677 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2685 1.287077e-02 -1.232629e-02 9.300469e-04 -1.265732e-01
#> Locus_2687 3.027631e-02 -1.490899e-02 7.625852e-03 -2.856393e-02
#> Locus_2696 5.416326e-02 -6.925806e-02 -2.517601e-02 -1.298426e-01
#> Locus_2698 1.467302e-01 1.898390e-02 4.119285e-02 1.068755e-01
#> Locus_2699 -5.333948e-02 3.796300e-02 -8.464196e-03 -1.042893e-01
#> Locus_2702 2.925599e-02 2.012611e-02 -7.514956e-03 -2.257755e-02
#> Locus_2703 5.662477e-02 1.357539e-02 -2.954128e-02 -9.550654e-02
#> Locus_2705 3.046626e-02 1.390273e-02 5.693283e-02 1.093393e-02
#> Locus_2712 6.171703e-02 2.167619e-02 -1.377394e-02 -9.406317e-02
#> Locus_2718 2.403273e-03 9.740097e-04 -2.217129e-02 -3.055069e-03
#> Locus_2719 4.621008e-02 1.891348e-02 2.981573e-02 -7.979274e-02
#> Locus_2724 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2725 -1.351362e-02 8.355912e-03 3.972568e-03 -3.699897e-02
#> Locus_2727 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2729 -6.654295e-02 -4.083687e-02 6.347781e-03 -2.901413e-02
#> Locus_2731 -5.466839e-03 1.467514e-02 -1.676098e-02 -5.148241e-02
#> Locus_2733 -1.395020e-01 -3.271483e-02 -1.379036e-02 2.078983e-02
#> Locus_2736 -1.302790e-02 -1.541389e-02 -2.709207e-03 1.858406e-02
#> Locus_2738 -4.777371e-02 -1.889937e-02 -1.316190e-02 1.240552e-02
#> Locus_2739 2.758581e-02 -5.851410e-02 -2.778175e-02 -4.692293e-02
#> Locus_2753 2.922095e-02 -3.230344e-02 1.243019e-03 -8.942868e-02
#> Locus_2755 1.154229e-02 -2.455218e-02 -8.606207e-04 -1.963150e-02
#> Locus_2758 2.375842e-01 6.646891e-03 4.273196e-02 1.598991e-01
#> Locus_2760 -5.907649e-04 8.595079e-02 -2.623208e-02 2.038197e-02
#> Locus_2761 -2.040637e-03 7.007019e-02 -3.028162e-02 9.251160e-03
#> Locus_2764 -4.777371e-02 -1.889937e-02 -1.316190e-02 1.240552e-02
#> Locus_2766 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2767 9.986514e-02 -8.337878e-02 -1.262851e-01 5.400032e-02
#> Locus_2768 -2.933256e-02 9.353876e-03 -1.567261e-02 2.844565e-03
#> Locus_2770 -5.527357e-02 2.646250e-02 2.742904e-03 -8.523613e-02
#> Locus_2774 -3.427026e-02 3.297567e-02 -1.311420e-02 -8.222833e-02
#> Locus_2776 3.510129e-02 7.240576e-02 -2.855259e-02 2.988994e-02
#> Locus_2789 1.446934e-01 -2.000225e-02 4.402679e-02 -2.953499e-01
#> Locus_2792 -1.094128e-02 -3.021668e-02 -4.352969e-03 2.887673e-02
#> Locus_2793 3.826741e-02 5.676546e-03 2.208693e-02 3.662925e-02
#> Locus_2794 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2795 5.294811e-04 4.200756e-02 -1.765412e-02 -8.652379e-02
#> Locus_2796 -8.522857e-03 -1.407305e-02 2.085590e-03 -2.034961e-03
#> Locus_2798 -3.817820e-02 -1.623095e-02 5.008129e-02 -1.343201e-02
#> Locus_2804 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2806 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2808 2.168856e-02 -2.092770e-02 4.121047e-03 2.644435e-02
#> Locus_2809 -1.906922e-02 4.987332e-03 4.650002e-03 -2.206098e-02
#> Locus_2810 -1.078759e-02 -5.129304e-02 -1.402064e-02 -1.286016e-02
#> Locus_2811 -2.981858e-02 -9.242478e-04 2.320969e-02 1.327416e-02
#> Locus_2814 -4.709292e-02 -1.244283e-02 2.670350e-03 -1.243836e-02
#> Locus_2816 -1.713513e-02 1.648784e-02 -6.557099e-03 -4.111417e-02
#> Locus_2817 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2818 -8.551230e-02 -6.578942e-04 -1.836721e-02 1.397094e-01
#> Locus_2823 1.441929e-01 -8.437236e-02 5.473632e-02 6.933347e-02
#> Locus_2830 -5.711269e-02 2.321144e-02 -6.246899e-03 -3.775928e-02
#> Locus_2831 2.475753e-01 1.257037e-01 7.691523e-02 -2.296390e-01
#> Locus_2832 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2835 2.315904e-02 -1.973334e-02 -2.187949e-02 2.832438e-02
#> Locus_2836 -3.331889e-02 -7.765895e-03 -4.616609e-03 2.890098e-02
#> Locus_2839 3.587350e-02 -1.571108e-01 -1.762882e-01 -3.750959e-03
#> Locus_2842 6.694382e-02 1.092691e-01 -6.771454e-02 -1.149399e-01
#> Locus_2844 1.055684e-02 -6.591640e-03 2.240391e-02 -6.818990e-02
#> Locus_2845 -2.789790e-02 -6.761920e-02 5.365551e-02 7.449406e-02
#> Locus_2850 -5.720765e-02 1.496199e-02 1.395001e-02 -5.883377e-02
#> Locus_2852 6.997756e-02 2.031820e-02 1.108053e-02 -8.981710e-02
#> Locus_2854 -9.432033e-03 1.683788e-03 1.964340e-03 1.823486e-02
#> Locus_2856 -4.082078e-02 1.234341e-01 2.419101e-02 -5.966975e-02
#> Locus_2861 4.237111e-02 1.282121e-02 4.486684e-02 6.339900e-02
#> Locus_2865 -2.437447e-02 -8.451602e-03 -9.773481e-03 1.105710e-02
#> Locus_2868 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2871 2.758581e-02 -5.851410e-02 -2.778175e-02 -4.692293e-02
#> Locus_2872 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2873 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2874 1.538092e-02 9.083490e-02 -2.304893e-02 -5.550717e-02
#> Locus_2875 1.883137e-02 -2.083875e-02 -8.113092e-03 -4.331455e-02
#> Locus_2878 1.254714e-02 -4.088695e-02 1.098845e-02 -1.587396e-02
#> Locus_2881 -1.146853e-02 -2.005932e-02 4.580551e-03 3.023669e-02
#> Locus_2884 3.136260e-03 2.393838e-02 -1.245169e-02 -3.138332e-02
#> Locus_2890 1.650051e-02 -2.758056e-02 -8.949828e-03 -3.721342e-02
#> Locus_2895 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2896 5.628349e-02 6.436711e-03 2.217141e-02 -4.134226e-02
#> Locus_2904 -9.657409e-02 1.865700e-02 1.409862e-02 1.524101e-01
#> Locus_2905 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2910 -9.961926e-03 7.022467e-03 3.802125e-02 1.104853e-02
#> Locus_2915 1.389105e-01 -2.055356e-03 -1.955714e-02 1.000609e-01
#> Locus_2916 1.892764e-02 -4.872766e-02 -5.781683e-02 3.502577e-02
#> Locus_2923 -2.380363e-02 -7.669923e-03 -1.289854e-02 4.396359e-02
#> Locus_2932 -1.486065e-02 8.728770e-02 3.688328e-02 -4.069814e-02
#> Locus_2933 2.244339e-02 -1.553036e-02 4.123681e-03 -3.956731e-02
#> Locus_2934 -4.943298e-02 7.264430e-03 -3.681829e-04 1.006709e-01
#> Locus_2936 -3.564173e-02 -5.656754e-02 7.034379e-02 -6.179412e-02
#> Locus_2937 1.356260e-01 6.297140e-02 2.388081e-02 -3.074054e-01
#> Locus_2938 -5.191095e-02 7.729676e-03 -9.836742e-03 4.742334e-02
#> Locus_2947 -2.627801e-01 -3.102543e-02 -5.494664e-02 1.980035e-01
#> Locus_2949 4.180244e-02 -2.767188e-02 -7.979945e-04 -5.460255e-02
#> Locus_2951 4.008027e-02 -1.057729e-02 -1.708614e-02 -5.392184e-02
#> Locus_2953 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2954 9.377496e-03 1.774501e-02 -1.874216e-03 2.588248e-02
#> Locus_2962 7.978111e-02 -2.459928e-02 -3.694813e-02 6.559458e-02
#> Locus_2966 -6.439166e-02 -4.131426e-02 3.528574e-02 -3.186445e-02
#> Locus_2972 1.616609e-01 -4.782868e-02 -3.569196e-02 1.395398e-01
#> Locus_2974 2.159870e-02 5.681731e-02 -8.974070e-04 -7.081833e-02
#> Locus_2976 -2.261760e-02 -7.985628e-03 -1.012200e-02 -1.431828e-02
#> Locus_2977 1.075386e-01 1.958617e-02 3.275403e-02 5.635043e-02
#> Locus_2978 -2.163496e-02 3.176507e-02 4.891780e-02 -1.214370e-01
#> Locus_2983 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2991 -3.136906e-02 -1.238923e-02 -1.305639e-02 1.484640e-02
#> Locus_2993 -1.004547e-01 -1.079328e-02 -3.010503e-02 1.325316e-01
#> Locus_2994 -2.310364e-02 1.368599e-02 2.809646e-02 -2.817464e-02
#> Locus_2995 -4.081275e-03 1.401404e-01 -6.056323e-02 1.850232e-02
#> Locus_3005 2.164795e-02 1.540099e-02 4.139541e-02 1.930629e-02
#> Locus_3009 -2.051130e-02 2.454332e-02 -2.033656e-02 4.582611e-02
#> Locus_3010 9.468429e-02 -1.097836e-02 -8.751551e-02 -2.442724e-01
#> Locus_3012 1.304420e-01 -6.344882e-02 -3.161698e-02 -3.058297e-01
#> Locus_3013 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3023 3.185957e-02 -2.198761e-02 3.994460e-02 4.470562e-02
#> Locus_3029 3.613703e-02 -9.229254e-03 1.247337e-02 -8.489260e-02
#> Locus_3030 5.167820e-02 -4.767546e-02 -4.664473e-02 5.961726e-02
#> Locus_3032 -4.681939e-02 2.117113e-03 -3.290217e-02 8.439566e-02
#> Locus_3042 1.593378e-02 3.382247e-02 2.218988e-02 -4.979344e-02
#> Locus_3043 2.656396e-02 8.882821e-02 1.232926e-02 -3.739770e-02
#> Locus_3046 -3.942022e-03 3.409632e-02 -3.859990e-04 1.526529e-02
#> Locus_3047 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3051 4.462584e-02 -2.471223e-02 6.068687e-02 -1.042301e-01
#> Locus_3054 -1.120851e-02 9.610688e-04 4.714717e-02 -9.722147e-03
#> Locus_3056 3.943557e-02 -3.206830e-02 -3.556916e-02 5.347639e-02
#> Locus_3058 8.808025e-02 -3.433903e-02 1.208423e-02 -9.824656e-02
#> Locus_3059 -1.438345e-02 -6.839072e-02 -1.869419e-02 -1.749200e-02
#> Locus_3060 3.861605e-02 -6.027085e-04 -1.096706e-04 2.766551e-02
#> Locus_3061 3.539335e-02 -9.972844e-04 -8.699050e-03 4.074271e-03
#> Locus_3062 -2.856486e-02 -7.765123e-02 2.438268e-02 7.998198e-02
#> Locus_3063 4.353563e-02 1.225073e-01 1.731918e-02 -8.629723e-02
#> Locus_3064 -3.060956e-03 1.051053e-01 -4.542242e-02 1.395131e-02
#> Locus_3069 1.003413e-01 1.938006e-02 -3.314920e-02 2.318880e-02
#> Locus_3078 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3079 1.593378e-02 3.382247e-02 2.218988e-02 -4.979344e-02
#> Locus_3081 1.883137e-02 -2.083875e-02 -8.113092e-03 -4.331455e-02
#> Locus_3084 3.564439e-02 3.450307e-02 5.239758e-04 -3.578198e-02
#> Locus_3087 -6.726868e-02 9.068811e-02 -3.606363e-02 -9.877367e-02
#> Locus_3089 -8.413628e-02 -5.303702e-02 1.927222e-02 6.529550e-02
#> Locus_3090 -1.400251e-02 -3.294148e-02 2.690253e-02 -2.012424e-02
#> Locus_3091 7.036151e-02 -6.753812e-02 -4.671610e-02 -2.890586e-02
#> Locus_3092 -4.757633e-02 -3.920838e-02 -4.161432e-03 1.940611e-02
#> Locus_3093 3.396953e-02 -2.829325e-02 -4.300166e-03 -5.831420e-02
#> Locus_3097 8.964341e-03 1.670232e-02 -3.241430e-02 -9.393061e-03
#> Locus_3098 3.955200e-02 -2.006721e-02 1.862973e-02 -1.124585e-01
#> Locus_3102 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3116 -3.620435e-02 2.147517e-02 -1.907097e-03 -6.317515e-02
#> Locus_3122 3.345068e-02 -4.094737e-02 8.420476e-02 -9.478300e-02
#> Locus_3126 5.807736e-02 3.007469e-02 8.116391e-02 4.999053e-02
#> Locus_3128 1.540176e-01 5.545472e-02 -9.935855e-03 -2.671859e-01
#> Locus_3129 2.560544e-02 6.973199e-03 1.907080e-02 2.271323e-02
#> Locus_3133 1.992385e-02 -1.404493e-02 -7.604249e-02 -2.209705e-02
#> Locus_3138 -4.931195e-02 -1.632999e-02 4.034819e-02 -1.019429e-02
#> Locus_3140 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3143 -1.544990e-01 -6.980656e-02 -3.215250e-02 -1.084453e-01
#> Locus_3145 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3147 2.807182e-02 -4.031463e-02 1.869425e-02 -6.640586e-02
#> Locus_3148 -5.361667e-02 2.146598e-02 -1.566517e-02 9.791606e-02
#> Locus_3154 2.895319e-02 7.325131e-03 4.170616e-02 -1.112905e-02
#> Locus_3155 1.082398e-02 7.700495e-03 2.069770e-02 1.186930e-02
#> Locus_3160 -1.250066e-01 6.898302e-03 -1.286209e-02 1.644658e-01
#> Locus_3161 4.545240e-02 -2.627559e-02 7.584742e-02 -8.734410e-03
#> Locus_3164 7.195639e-02 9.513053e-02 -7.929670e-02 -8.886431e-02
#> Locus_3165 4.255709e-03 -3.054220e-02 -2.608780e-02 -1.993497e-02
#> Locus_3167 2.169192e-02 -1.708223e-02 2.186961e-03 -1.057947e-03
#> Locus_3168 -1.070327e-02 9.166716e-03 2.033171e-02 -2.100497e-02
#> Locus_3169 -7.380745e-02 -1.187172e-03 -1.014166e-02 1.163163e-01
#> Locus_3171 2.972480e-02 4.511996e-04 4.258554e-02 5.079633e-02
#> Locus_3175 6.922020e-02 -5.801991e-02 2.799706e-03 -1.054883e-01
#> Locus_3177 -8.123662e-02 -3.134028e-02 2.772410e-02 -1.570040e-01
#> Locus_3179 -7.771002e-02 1.528943e-02 1.016994e-02 1.251169e-01
#> Locus_3181 3.627037e-02 -1.813464e-02 -1.604563e-02 -6.806492e-03
#> Locus_3182 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3188 -1.720300e-02 -7.796293e-02 8.249405e-02 -2.784405e-01
#> Locus_3190 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3191 1.409170e-03 -1.368952e-02 -1.250036e-02 1.636349e-02
#> Locus_3192 -6.485476e-02 -5.012819e-03 -9.031063e-03 4.867145e-02
#> Locus_3197 -7.038246e-02 1.268720e-01 -5.106461e-02 1.085761e-01
#> Locus_3198 -2.340970e-02 1.058556e-03 -1.645108e-02 3.760957e-02
#> PC2 PC3
#> Locus_10 3.045178e-02 -5.346571e-03
#> Locus_15 3.932804e-02 -1.482929e-01
#> Locus_22 -1.622119e-02 -5.073074e-02
#> Locus_28 3.835175e-02 1.033904e-02
#> Locus_32 2.562468e-02 1.819505e-02
#> Locus_35 -1.621445e-16 6.195147e-17
#> Locus_37 1.024925e-16 -4.830201e-17
#> Locus_61 -1.887042e-01 1.138077e-01
#> Locus_71 -9.594437e-03 -3.834203e-02
#> Locus_72 6.383109e-17 4.001075e-17
#> Locus_82 -2.594515e-17 1.071368e-16
#> Locus_84 8.254975e-03 -8.667521e-03
#> Locus_90 1.492710e-01 9.758328e-02
#> Locus_91 1.085666e-01 -1.456370e-02
#> Locus_96 -4.663135e-19 -3.993036e-17
#> Locus_104 2.642961e-03 7.188306e-03
#> Locus_122 -4.669036e-02 1.004358e-02
#> Locus_125 -5.162213e-02 -5.808578e-02
#> Locus_131 -1.227597e-02 -1.996908e-02
#> Locus_133 -3.716814e-17 4.312769e-18
#> Locus_138 -1.186612e-01 1.378807e-01
#> Locus_154 -7.059888e-02 -1.073117e-01
#> Locus_162 -2.811967e-02 -5.802721e-02
#> Locus_166 1.584043e-01 3.682674e-03
#> Locus_177 -4.046870e-17 2.625603e-17
#> Locus_185 4.435703e-17 -2.805914e-18
#> Locus_191 2.113801e-02 4.893365e-02
#> Locus_193 -1.328142e-02 -2.049286e-02
#> Locus_207 -9.760745e-02 7.933000e-02
#> Locus_211 4.302135e-02 -3.553544e-03
#> Locus_213 1.153558e-02 4.292904e-02
#> Locus_214 9.960284e-02 -1.087082e-02
#> Locus_222 1.421824e-02 8.747202e-03
#> Locus_227 -3.169478e-02 -1.587278e-01
#> Locus_242 2.206421e-02 7.332887e-02
#> Locus_249 -1.775299e-01 1.570664e-01
#> Locus_254 -1.747252e-02 5.211207e-02
#> Locus_256 -1.397581e-01 1.567349e-01
#> Locus_263 2.557354e-01 1.605037e-01
#> Locus_268 4.257690e-03 2.393699e-02
#> Locus_279 5.556950e-02 1.691014e-02
#> Locus_285 -3.627584e-02 4.293768e-02
#> Locus_287 -6.040910e-02 -4.250385e-02
#> Locus_292 -1.278210e-02 -4.187395e-03
#> Locus_294 -1.098070e-02 1.078969e-02
#> Locus_303 -2.589549e-17 1.974474e-17
#> Locus_308 2.569663e-02 -3.483250e-02
#> Locus_312 4.957827e-03 5.953194e-02
#> Locus_313 -1.291429e-17 -2.510441e-17
#> Locus_314 3.510152e-02 -4.550986e-02
#> Locus_317 -2.816775e-02 -2.508352e-02
#> Locus_318 -1.648631e-01 4.442576e-02
#> Locus_323 0.000000e+00 0.000000e+00
#> Locus_330 -5.155683e-03 -8.809137e-03
#> Locus_331 -1.925404e-02 3.741467e-03
#> Locus_333 2.681612e-02 -2.100526e-03
#> Locus_334 1.253263e-02 1.825983e-02
#> Locus_336 -3.671942e-02 -2.576085e-02
#> Locus_337 3.173298e-02 2.877134e-02
#> Locus_339 5.311015e-03 -2.564898e-02
#> Locus_341 4.591717e-02 -3.130790e-02
#> Locus_345 -2.996330e-04 -1.385930e-02
#> Locus_350 -1.760069e-02 4.918540e-02
#> Locus_354 -1.496875e-02 -4.443105e-03
#> Locus_359 5.870727e-02 -1.425911e-01
#> Locus_365 -4.441294e-03 -3.008199e-02
#> Locus_366 -1.146315e-02 -1.345989e-02
#> Locus_369 -2.953801e-02 1.437419e-02
#> Locus_372 -9.832016e-03 -1.827062e-02
#> Locus_389 5.686505e-02 6.981765e-03
#> Locus_391 4.899027e-02 -1.025690e-02
#> Locus_392 1.425629e-01 -1.474227e-02
#> Locus_393 1.035286e-02 1.570528e-03
#> Locus_394 0.000000e+00 0.000000e+00
#> Locus_396 -1.405258e-02 -4.208111e-02
#> Locus_399 9.652079e-02 -1.817654e-02
#> Locus_400 3.765233e-03 1.413981e-02
#> Locus_401 1.653627e-02 -5.447054e-02
#> Locus_404 4.305983e-03 -2.048100e-01
#> Locus_406 -1.932925e-01 -5.372249e-02
#> Locus_409 -1.123084e-02 -3.248241e-02
#> Locus_420 -2.278934e-02 -2.105183e-02
#> Locus_422 2.379583e-02 -1.090216e-02
#> Locus_427 -1.598734e-02 -1.086197e-01
#> Locus_430 0.000000e+00 0.000000e+00
#> Locus_436 3.533039e-02 -2.923329e-03
#> Locus_438 -3.324724e-03 4.212286e-02
#> Locus_439 1.981725e-02 -8.056889e-03
#> Locus_441 -1.008897e-01 1.539214e-01
#> Locus_445 -6.825160e-03 -1.069402e-02
#> Locus_448 -2.208951e-02 -3.195129e-02
#> Locus_449 -1.407264e-03 -9.903051e-02
#> Locus_456 -4.461788e-03 5.110399e-03
#> Locus_457 -6.726335e-04 -2.094037e-02
#> Locus_458 -6.058657e-02 2.975126e-02
#> Locus_460 0.000000e+00 0.000000e+00
#> Locus_461 1.384492e-02 -1.136256e-02
#> Locus_462 0.000000e+00 0.000000e+00
#> Locus_464 4.764153e-03 -1.901145e-02
#> Locus_469 1.776919e-01 -6.198690e-02
#> Locus_475 2.887923e-03 6.425220e-02
#> Locus_480 1.866194e-01 -1.204741e-02
#> Locus_482 -2.552886e-02 7.208853e-02
#> Locus_483 6.385891e-02 2.193880e-02
#> Locus_491 -2.400052e-02 1.786822e-02
#> Locus_498 -1.186297e-02 -5.084047e-02
#> Locus_500 -3.153273e-03 3.813358e-03
#> Locus_503 -2.790321e-02 -8.876553e-02
#> Locus_505 3.801522e-02 -1.188309e-01
#> Locus_508 -1.026807e-01 -1.817271e-01
#> Locus_512 -4.113461e-02 -3.988587e-02
#> Locus_516 1.636917e-02 1.379109e-02
#> Locus_517 1.511321e-03 8.666238e-02
#> Locus_518 -2.430397e-02 -4.587874e-02
#> Locus_528 0.000000e+00 0.000000e+00
#> Locus_529 -6.148224e-02 7.295088e-02
#> Locus_531 1.094699e-01 -1.967392e-02
#> Locus_532 3.561819e-02 -3.025294e-02
#> Locus_533 -3.329951e-02 -3.484937e-03
#> Locus_535 2.258525e-02 -2.228672e-02
#> Locus_546 3.304563e-02 -9.415872e-02
#> Locus_557 0.000000e+00 0.000000e+00
#> Locus_558 -8.485978e-02 7.104232e-03
#> Locus_561 0.000000e+00 0.000000e+00
#> Locus_562 1.622569e-02 1.730259e-02
#> Locus_564 -1.906256e-02 -5.622436e-02
#> Locus_566 3.850902e-03 2.520877e-02
#> Locus_572 4.112960e-04 -6.982841e-03
#> Locus_573 6.660270e-03 1.579557e-02
#> Locus_579 2.237378e-02 -2.851244e-02
#> Locus_581 -1.812083e-02 6.496368e-02
#> Locus_582 6.660270e-03 1.579557e-02
#> Locus_583 7.426727e-02 1.459449e-02
#> Locus_586 5.349295e-03 5.522988e-02
#> Locus_589 -8.388258e-02 1.507841e-01
#> Locus_596 -4.153703e-03 -1.632009e-02
#> Locus_597 -3.371831e-02 -4.218834e-02
#> Locus_598 0.000000e+00 0.000000e+00
#> Locus_600 -1.607689e-02 -1.352479e-02
#> Locus_602 1.267743e-02 6.724328e-02
#> Locus_603 0.000000e+00 0.000000e+00
#> Locus_604 -9.409582e-06 1.486332e-02
#> Locus_609 7.737600e-02 7.780757e-02
#> Locus_612 2.438375e-02 1.412396e-01
#> Locus_613 4.356745e-02 2.889999e-02
#> Locus_614 -1.161941e-02 -2.157992e-02
#> Locus_622 -9.231510e-03 1.068113e-02
#> Locus_625 -1.126233e-02 2.788164e-02
#> Locus_626 -4.800103e-02 3.573644e-02
#> Locus_633 -8.572524e-05 1.915475e-02
#> Locus_634 -3.074112e-02 3.647544e-02
#> Locus_635 1.175244e-01 2.783722e-02
#> Locus_636 2.237551e-01 4.536555e-02
#> Locus_637 -5.338427e-02 5.884299e-02
#> Locus_639 -7.243818e-03 -2.276881e-03
#> Locus_642 0.000000e+00 0.000000e+00
#> Locus_647 -6.651639e-02 -5.720298e-02
#> Locus_652 -3.158542e-02 -2.390446e-02
#> Locus_665 1.190078e-02 2.089840e-03
#> Locus_671 -8.375718e-02 -6.366363e-02
#> Locus_679 -7.512514e-03 -5.058438e-02
#> Locus_681 -7.298408e-02 2.838458e-03
#> Locus_682 -8.375718e-02 -6.366363e-02
#> Locus_683 0.000000e+00 0.000000e+00
#> Locus_685 6.660270e-03 1.579557e-02
#> Locus_687 -3.817671e-03 -9.222976e-03
#> Locus_691 -1.783893e-02 -2.399689e-02
#> Locus_699 1.493229e-01 1.254813e-02
#> Locus_701 0.000000e+00 0.000000e+00
#> Locus_707 8.498023e-02 7.939972e-03
#> Locus_710 6.554793e-02 3.922504e-02
#> Locus_711 5.878413e-03 2.726093e-02
#> Locus_712 0.000000e+00 0.000000e+00
#> Locus_717 -7.512514e-03 -5.058438e-02
#> Locus_718 0.000000e+00 0.000000e+00
#> Locus_724 1.466928e-03 -2.532591e-02
#> Locus_725 -3.074112e-02 3.647544e-02
#> Locus_727 2.977048e-02 -2.523673e-03
#> Locus_729 0.000000e+00 0.000000e+00
#> Locus_733 -1.141271e-01 4.976420e-02
#> Locus_734 -2.065568e-03 -2.285235e-02
#> Locus_735 -6.192609e-02 1.071008e-01
#> Locus_736 -8.375718e-02 -6.366363e-02
#> Locus_740 1.077737e-02 -1.254852e-01
#> Locus_742 8.345158e-03 -3.809343e-02
#> Locus_746 6.357169e-02 -3.321041e-02
#> Locus_750 6.922461e-03 -5.681279e-03
#> Locus_751 -1.846302e-02 2.136226e-02
#> Locus_753 1.099579e-02 1.523274e-02
#> Locus_758 -2.264315e-02 2.236755e-02
#> Locus_765 -3.114670e-02 2.393619e-02
#> Locus_769 -2.378532e-02 4.038944e-02
#> Locus_770 3.305054e-01 -5.486374e-02
#> Locus_771 -1.796101e-01 -7.116739e-02
#> Locus_776 0.000000e+00 0.000000e+00
#> Locus_779 4.432321e-03 6.759983e-03
#> Locus_784 -1.367169e-02 1.895486e-02
#> Locus_786 -1.040577e-01 4.682593e-03
#> Locus_787 -6.130818e-03 1.325471e-02
#> Locus_798 -1.029294e-02 -1.529328e-01
#> Locus_802 -7.041089e-02 -1.278405e-01
#> Locus_803 -2.408085e-02 5.227100e-02
#> Locus_807 1.384492e-02 -1.136256e-02
#> Locus_808 -2.232526e-02 -1.676092e-02
#> Locus_809 -1.846302e-02 2.136226e-02
#> Locus_811 -6.432863e-03 -1.283299e-02
#> Locus_816 -5.055433e-03 5.781392e-03
#> Locus_821 -2.002410e-03 -1.262250e-02
#> Locus_825 -2.888971e-02 -9.765034e-02
#> Locus_831 2.579111e-02 1.880962e-02
#> Locus_835 -9.334739e-02 2.754044e-02
#> Locus_838 -1.152508e-01 -3.843951e-02
#> Locus_847 -1.451603e-02 2.037775e-02
#> Locus_850 -4.501039e-03 2.698836e-02
#> Locus_851 6.685996e-02 -1.487041e-02
#> Locus_853 0.000000e+00 0.000000e+00
#> Locus_854 1.900761e-02 -5.941547e-02
#> Locus_859 -5.162213e-02 -5.808578e-02
#> Locus_861 2.451783e-03 1.253622e-02
#> Locus_864 1.693219e-01 -5.532680e-02
#> Locus_869 -9.678693e-02 -7.268148e-02
#> Locus_870 2.024300e-02 2.590985e-02
#> Locus_875 0.000000e+00 0.000000e+00
#> Locus_876 2.161092e-02 -6.193747e-02
#> Locus_879 -1.833613e-02 -2.491593e-02
#> Locus_881 2.715637e-01 -4.648512e-02
#> Locus_887 2.611969e-02 -5.673754e-03
#> Locus_888 0.000000e+00 0.000000e+00
#> Locus_895 3.643582e-02 -5.653798e-03
#> Locus_896 1.873458e-01 -2.554388e-02
#> Locus_897 -9.453363e-03 -1.666201e-02
#> Locus_898 -4.069194e-02 6.320078e-02
#> Locus_899 -8.118094e-02 -1.820211e-02
#> Locus_901 -6.127364e-02 -6.388458e-02
#> Locus_902 -1.932084e-01 -5.981175e-02
#> Locus_903 3.925061e-02 -3.350561e-02
#> Locus_905 3.118422e-02 3.860608e-02
#> Locus_908 0.000000e+00 0.000000e+00
#> Locus_909 -4.445926e-02 -5.113949e-02
#> Locus_915 0.000000e+00 0.000000e+00
#> Locus_924 3.450165e-02 3.781232e-02
#> Locus_932 2.203238e-02 1.089852e-02
#> Locus_935 0.000000e+00 0.000000e+00
#> Locus_936 4.876120e-02 -7.593039e-02
#> Locus_938 -6.148224e-02 7.295088e-02
#> Locus_941 0.000000e+00 0.000000e+00
#> Locus_944 5.075236e-02 2.234779e-02
#> Locus_945 -2.207664e-01 -3.677252e-02
#> Locus_946 0.000000e+00 0.000000e+00
#> Locus_949 1.421023e-02 7.483762e-03
#> Locus_950 1.341497e-02 -4.732179e-02
#> Locus_951 2.509717e-01 -3.051882e-01
#> Locus_953 2.577630e-02 -2.144126e-02
#> Locus_958 0.000000e+00 0.000000e+00
#> Locus_965 1.629759e-02 -1.353872e-02
#> Locus_966 7.572146e-02 -9.820120e-03
#> Locus_967 4.968669e-02 -6.683669e-03
#> Locus_968 5.604291e-02 3.166758e-02
#> Locus_970 -2.647040e-02 1.996320e-02
#> Locus_972 -1.879633e-02 -3.088597e-02
#> Locus_974 1.358273e-02 1.011429e-02
#> Locus_979 4.280257e-03 -5.447616e-03
#> Locus_992 -1.137742e-02 -3.261464e-02
#> Locus_998 -2.252466e-02 5.576328e-02
#> Locus_999 1.978228e-02 -2.160919e-02
#> Locus_1002 -2.154583e-02 3.413531e-02
#> Locus_1005 2.419613e-01 -6.135268e-02
#> Locus_1006 -4.010118e-03 -5.063028e-02
#> Locus_1007 7.761012e-02 -5.981956e-03
#> Locus_1015 8.501101e-02 -2.412346e-02
#> Locus_1016 0.000000e+00 0.000000e+00
#> Locus_1017 -6.903259e-02 -5.608564e-02
#> Locus_1023 -3.074112e-02 3.647544e-02
#> Locus_1024 1.967148e-03 1.254086e-02
#> Locus_1025 2.072573e-02 5.065099e-02
#> Locus_1027 -1.048752e-02 -1.819992e-02
#> Locus_1030 2.112239e-01 -8.353155e-02
#> Locus_1031 7.322983e-02 4.671516e-02
#> Locus_1033 -5.060739e-02 -4.051436e-02
#> Locus_1034 -1.784095e-02 -3.181419e-02
#> Locus_1035 2.673119e-02 4.367773e-04
#> Locus_1038 -9.409582e-06 1.486332e-02
#> Locus_1041 8.076017e-03 -4.025031e-02
#> Locus_1042 3.079678e-02 1.904378e-02
#> Locus_1044 3.856180e-02 -1.352527e-01
#> Locus_1046 -1.532355e-02 -1.084297e-02
#> Locus_1051 -2.581107e-02 -2.904289e-02
#> Locus_1054 -6.224494e-03 -1.668903e-02
#> Locus_1055 0.000000e+00 0.000000e+00
#> Locus_1065 9.117260e-02 5.742010e-02
#> Locus_1071 -4.528629e-02 4.473510e-02
#> Locus_1072 -3.188299e-02 -4.392195e-02
#> Locus_1073 -3.612656e-02 3.224362e-02
#> Locus_1074 4.103242e-02 -6.251980e-03
#> Locus_1075 2.735085e-02 -2.494841e-02
#> Locus_1076 0.000000e+00 0.000000e+00
#> Locus_1080 -4.999517e-02 4.021691e-02
#> Locus_1082 0.000000e+00 0.000000e+00
#> Locus_1093 -6.455430e-03 1.655161e-02
#> Locus_1095 0.000000e+00 0.000000e+00
#> Locus_1098 -2.378532e-02 4.038944e-02
#> Locus_1099 -3.671942e-02 -2.576085e-02
#> Locus_1105 -1.024317e-01 9.859239e-02
#> Locus_1106 2.308714e-02 -5.624762e-03
#> Locus_1107 -9.553846e-03 1.272426e-03
#> Locus_1108 1.332054e-02 3.159113e-02
#> Locus_1109 0.000000e+00 0.000000e+00
#> Locus_1113 -2.536910e-02 -4.859764e-02
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#> Locus_1144 6.512970e-03 3.525148e-02
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#> Locus_1162 1.146094e-01 1.302881e-02
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#> Locus_1191 -5.925267e-03 5.008297e-03
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#> Locus_1201 1.145791e-02 -1.994507e-01
#> Locus_1204 4.257690e-03 2.393699e-02
#> Locus_1206 -1.367169e-02 1.895486e-02
#> Locus_1211 1.668752e-02 -4.789968e-03
#> Locus_1212 -2.135061e-02 9.435121e-03
#> Locus_1217 -1.455580e-02 3.428537e-02
#> Locus_1223 -2.208951e-02 -3.195129e-02
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#> Locus_1231 1.332054e-02 3.159113e-02
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#> Locus_1254 1.358273e-02 1.011429e-02
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#> Locus_1256 -4.684839e-02 6.345304e-02
#> Locus_1258 -8.652554e-02 1.435958e-01
#> Locus_1262 1.015304e-01 -8.490285e-03
#> Locus_1263 1.193775e-02 8.418795e-02
#> Locus_1265 -5.972884e-03 -1.787988e-02
#> Locus_1266 9.776103e-03 -4.873434e-02
#> Locus_1270 9.777093e-03 -3.791863e-02
#> Locus_1278 -2.438352e-01 -2.897916e-02
#> Locus_1279 -7.740777e-02 1.198426e-02
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#> Locus_1287 -4.170971e-02 -9.289683e-02
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#> Locus_1305 -1.948588e-02 -8.429359e-03
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#> Locus_1319 2.716546e-02 2.022857e-02
#> Locus_1338 5.324282e-02 -9.487404e-02
#> Locus_1343 7.226036e-02 2.501653e-04
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#> Locus_1364 1.341099e-02 3.177524e-02
#> Locus_1366 -8.520344e-03 1.977547e-02
#> Locus_1370 -1.022264e-01 -6.302124e-02
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#> Locus_1377 -1.290924e-02 -3.857397e-02
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#> Locus_1384 -3.034029e-02 8.892583e-03
#> Locus_1392 1.312295e-01 2.203025e-02
#> Locus_1396 4.868682e-02 -2.548322e-01
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#> Locus_1414 -8.792007e-02 1.014053e-01
#> Locus_1416 5.137571e-02 1.060257e-02
#> Locus_1420 5.869623e-02 -2.833525e-02
#> Locus_1423 -7.001448e-02 -1.000199e-01
#> Locus_1424 7.087683e-02 -9.241908e-02
#> Locus_1425 -1.261401e-02 2.975968e-03
#> Locus_1426 1.812007e-01 1.055684e-01
#> Locus_1427 -9.231510e-03 1.068113e-02
#> Locus_1430 -7.937818e-02 -1.404631e-01
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#> Locus_1439 -7.229283e-02 -1.591074e-02
#> Locus_1447 5.595269e-03 -1.204074e-01
#> Locus_1452 -8.786929e-02 -6.573245e-02
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#> Locus_1460 -2.861474e-02 1.968198e-02
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#> Locus_1468 -1.084933e-01 -3.137657e-02
#> Locus_1470 -1.278145e-02 1.116422e-01
#> Locus_1477 1.264703e-01 -6.845092e-02
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#> Locus_1482 4.343415e-03 4.782237e-03
#> Locus_1485 -4.501039e-03 2.698836e-02
#> Locus_1486 -2.416652e-02 -2.563312e-02
#> Locus_1487 -2.953984e-02 -4.418587e-02
#> Locus_1488 -1.315208e-02 7.576682e-03
#> Locus_1489 -5.155683e-03 -8.809137e-03
#> Locus_1490 -1.548992e-02 -1.198679e-02
#> Locus_1494 -2.102438e-02 -2.923239e-02
#> Locus_1496 7.274588e-03 -4.782299e-02
#> Locus_1500 2.294062e-03 2.275316e-02
#> Locus_1502 2.727592e-02 -1.216356e-02
#> Locus_1506 4.682010e-03 -1.568052e-02
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#> Locus_1509 2.553234e-02 2.316354e-02
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#> Locus_1598 -6.432863e-03 -1.283299e-02
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#> Locus_1602 -5.925267e-03 5.008297e-03
#> Locus_1603 4.257690e-03 2.393699e-02
#> Locus_1604 8.916690e-04 -1.252419e-01
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#> Locus_1612 7.475324e-03 -3.886652e-02
#> Locus_1614 -6.040910e-02 -4.250385e-02
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#> Locus_1661 -3.734406e-02 1.687137e-03
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#> Locus_1708 -5.166114e-03 6.958573e-03
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#> Locus_1870 -5.925267e-03 5.008297e-03
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#> Locus_1948 -1.290924e-02 -3.857397e-02
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#> Locus_1958 4.889312e-02 -6.169322e-03
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#> Locus_1962 -1.391049e-01 1.292474e-01
#> Locus_1965 6.393015e-02 5.999470e-03
#> Locus_1967 1.332054e-02 3.159113e-02
#> Locus_1968 2.232475e-02 -1.448173e-02
#> Locus_1971 1.593190e-02 6.429880e-03
#> Locus_1974 4.049149e-02 1.913298e-02
#> Locus_1977 -6.223051e-02 4.529438e-02
#> Locus_1982 -5.329960e-02 1.841953e-03
#> Locus_1983 2.942575e-02 -6.636752e-02
#> Locus_1984 -7.886172e-03 -6.119198e-03
#> Locus_1988 9.565422e-03 1.507026e-03
#> Locus_1990 -1.912664e-02 -1.001321e-01
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#> Locus_1993 0.000000e+00 0.000000e+00
#> Locus_2003 -2.836043e-03 -5.648479e-02
#> Locus_2007 1.332054e-02 3.159113e-02
#> Locus_2014 -4.113461e-02 -3.988587e-02
#> Locus_2016 -4.800103e-02 3.573644e-02
#> Locus_2017 6.328590e-02 -5.003563e-02
#> Locus_2019 2.664221e-02 -3.969144e-03
#> Locus_2020 9.429616e-03 1.178236e-02
#> Locus_2021 4.112772e-02 1.320519e-02
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#> Locus_2060 -3.850809e-02 7.482934e-03
#> Locus_2063 2.642961e-03 7.188306e-03
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#> Locus_3098 1.601821e-02 -6.855016e-03
#> Locus_3102 0.000000e+00 0.000000e+00
#> Locus_3116 1.358273e-02 1.011429e-02
#> Locus_3122 2.588005e-01 -1.735688e-02
#> Locus_3126 6.862016e-02 -2.838669e-02
#> Locus_3128 -4.223499e-02 3.779781e-02
#> Locus_3129 3.467066e-02 -1.074685e-02
#> Locus_3133 -3.021985e-02 -3.315961e-02
#> Locus_3138 1.175861e-02 -8.921509e-02
#> Locus_3140 0.000000e+00 0.000000e+00
#> Locus_3143 -1.679793e-01 2.471014e-01
#> Locus_3145 0.000000e+00 0.000000e+00
#> Locus_3147 -9.984608e-02 1.120047e-01
#> Locus_3148 -9.353263e-02 -4.156787e-02
#> Locus_3154 8.186835e-03 -1.129009e-02
#> Locus_3155 2.276988e-02 -1.283669e-02
#> Locus_3160 -1.831819e-01 -6.600054e-02
#> Locus_3161 -5.323225e-02 7.189063e-02
#> Locus_3164 2.294497e-02 3.974707e-02
#> Locus_3165 -3.156946e-03 -1.347636e-03
#> Locus_3167 -1.437195e-02 5.026653e-02
#> Locus_3168 1.516292e-02 -2.898269e-02
#> Locus_3169 -1.505058e-01 -3.107451e-02
#> Locus_3171 2.996545e-02 2.829648e-02
#> Locus_3175 -3.005342e-02 -3.695550e-02
#> Locus_3177 -6.443338e-02 4.103366e-02
#> Locus_3179 -1.420899e-01 -7.208901e-02
#> Locus_3181 1.471643e-02 -8.001551e-04
#> Locus_3182 0.000000e+00 0.000000e+00
#> Locus_3188 7.203400e-02 -1.238268e-01
#> Locus_3190 0.000000e+00 0.000000e+00
#> Locus_3191 5.557947e-02 -1.193839e-03
#> Locus_3192 2.388018e-02 -7.847234e-03
#> Locus_3197 -1.063482e-01 -3.514916e-02
#> Locus_3198 -4.669617e-02 -4.231957e-02
#>
#> $sites
#> RDA1 RDA2 RDA3 PC1 PC2
#> ALT3 -0.607358319 1.58563933 0.3496179 -3.55018546 0.5751105753
#> BAR360 1.902849544 -0.76604821 -0.6439038 -2.12447635 -0.2722833852
#> BLL5 0.693520569 1.96873515 -2.1864553 -0.81108561 -1.6625760009
#> BNT5 -2.615784176 -0.29205461 -1.0298086 0.53560509 -0.7971357244
#> BOF1 1.590877723 -0.21821229 0.1445185 0.96823582 0.7194896725
#> BOT3 -1.136573356 -1.45182344 -1.1999707 -0.12372612 -0.2651981743
#> CAR1 -0.667995445 -0.21186827 0.3931249 1.03635154 -0.0008125122
#> CTR7 2.201377522 -1.32249087 -0.9910732 1.53673955 1.5430384989
#> CYR26 -0.955298448 0.56643109 -0.5141383 0.24562661 1.2458162395
#> FCT13 2.032409896 2.45540518 0.9478023 -2.15437994 -0.8175669328
#> HAY2 -1.329515351 -0.74983461 -0.3072347 -0.83641880 0.6454900542
#> HHR2 2.687823979 -1.00040226 -0.6553835 1.42066544 2.6872436304
#> HNT18 -2.885862993 -0.14606260 0.3243279 1.42193003 -1.3638930576
#> HOP1 0.574089901 -1.31523313 1.5303251 -1.01928580 -2.6544788183
#> INK1 2.073849506 0.64761325 0.8921924 0.49979492 -0.9055914561
#> INK6 1.732176587 1.10036647 0.7741716 0.40724720 -1.3231796846
#> KBG6 0.165409301 0.90007607 1.9804961 0.37907290 1.5493482879
#> KRV3 -0.650153922 0.21324052 3.1608084 0.95403400 1.3047334526
#> LBH40 0.314716732 3.14855253 -0.9762156 -0.52049296 0.7182661429
#> LCV19 -3.024180882 -0.89616379 -0.7052436 0.95477464 -1.2752955004
#> LFV10 0.521554662 -1.80899799 -2.7789950 0.09156243 1.6031745397
#> LTF1 -0.791869749 1.04431662 0.4428249 -1.90495327 0.5977506625
#> LTF6 -0.881138255 0.71503254 0.3282726 -1.58765481 0.2282181833
#> MLG2 1.006613722 -1.97252203 -0.3702433 -0.85647358 0.3750514845
#> MLG5 1.365806253 -2.57632659 -1.3250168 -1.59764781 -0.0074023270
#> MNR2 -0.881848061 -0.76578848 1.7464003 -0.57992342 1.1094446997
#> MRO3 0.910221010 -0.05504183 -0.6512574 0.92793262 1.4134687641
#> OLA14 -2.571601093 -0.15835480 0.1138543 1.57457009 -1.8896490327
#> OLA9 -3.051388597 0.17810191 -0.3897725 1.95997884 -3.1707018189
#> POT1 1.154826338 -1.61435523 -2.9364790 0.38807069 2.8330896133
#> PRU1 -1.033656848 1.31800096 1.2212348 0.91229411 -0.4608526034
#> PRU6 -0.650441272 1.45812500 1.6221330 0.68651209 -0.5116435167
#> PT8 -1.449168211 -2.75502228 3.5610375 -0.12200635 0.9958066956
#> REY5 -1.462480526 -0.33453217 0.2292414 1.24674474 0.1156194052
#> RPS38 -2.814964598 -0.07399529 -1.1845029 2.01997659 -1.6000988948
#> RPS9 -2.706653253 -0.19986664 -0.9469557 1.62378263 -2.0160940939
#> SBR10 -1.286467540 0.78920473 -1.3592337 1.33384428 0.0947524518
#> SKY3 -0.839269641 -1.87395237 -0.6689273 -0.39995703 -0.3835032518
#> SKY5 -0.851538504 -1.66104556 -1.9641636 -0.35525520 -0.7312027157
#> SRM1 1.524051685 -0.87422127 0.4490775 -1.61571066 -0.1729069906
#> SRM7 1.679672853 -0.81725041 0.5250890 -2.24534752 -0.0054536726
#> SUT7 -0.008857422 1.19205853 1.9012667 -0.84520937 -1.0269243787
#> TTC1 1.385196284 -0.10004517 -0.1863376 0.93636800 1.0276256196
#> TTC2 1.597050779 0.01714006 0.2199844 0.98248003 1.2805980822
#> WAW1 1.279866651 0.21646449 -0.1649088 1.24710313 0.6633282699
#> WDC1 0.694610060 0.54436960 1.4008303 0.64218008 0.3561911663
#> WDC2 2.002688772 -0.48912003 1.5356288 1.02490718 1.9661662864
#> WHR2 0.305374655 -0.76345450 -0.9441282 -0.89529210 -1.9552230969
#> WTW14 1.316097160 -0.45564281 -0.1354000 -1.29521166 -1.2772384337
#> WTW3 1.810770179 -0.52792069 0.4112121 -1.17126810 -0.4047634964
#> YUC3 1.405337484 0.01604836 2.4897832 -2.14525585 1.1662467386
#> ZUM18 -0.502297042 4.47697976 -2.0077235 0.40585515 0.0614906320
#> ZUM20 -0.272476303 3.69574808 -1.4717833 0.39297734 0.0751097215
#> PC3
#> ALT3 1.36393839
#> BAR360 0.32928135
#> BLL5 0.32307359
#> BNT5 0.92230946
#> BOF1 0.00126034
#> BOT3 -1.77036971
#> CAR1 1.28343963
#> CTR7 -0.39837430
#> CYR26 0.45356677
#> FCT13 -1.54412883
#> HAY2 -3.35610236
#> HHR2 -0.48730308
#> HNT18 0.03353609
#> HOP1 3.14963379
#> INK1 -1.57155262
#> INK6 -0.93628485
#> KBG6 0.92436500
#> KRV3 1.43165679
#> LBH40 0.13490647
#> LCV19 0.62060155
#> LFV10 -0.08982798
#> LTF1 -0.49057531
#> LTF6 0.62070618
#> MLG2 0.41294351
#> MLG5 1.65400213
#> MNR2 -1.50388330
#> MRO3 1.19085326
#> OLA14 1.03326291
#> OLA9 -2.22443495
#> POT1 1.00535396
#> PRU1 1.97510046
#> PRU6 0.43246364
#> PT8 -1.77439014
#> REY5 1.21030440
#> RPS38 -0.92152389
#> RPS9 -1.82713565
#> SBR10 1.01613971
#> SKY3 -2.59756340
#> SKY5 -0.73327785
#> SRM1 -1.08994543
#> SRM7 -0.88334213
#> SUT7 1.74380282
#> TTC1 0.18045650
#> TTC2 -0.14274102
#> WAW1 0.64676783
#> WDC1 -0.04577562
#> WDC2 -1.10844126
#> WHR2 1.93142556
#> WTW14 -0.48709556
#> WTW3 -0.86315517
#> YUC3 -0.19366717
#> ZUM18 0.94898317
#> ZUM20 0.06675630
#>
#> $constraints
#> RDA1 RDA2 RDA3 PC1 PC2
#> ALT3 -1.47960907 1.42371542 -0.56620185 -3.55018546 0.5751105753
#> BAR360 0.99527548 -1.10206846 -0.72739449 -2.12447635 -0.2722833852
#> BLL5 0.77406582 1.44223623 -2.79895634 -0.81108561 -1.6625760009
#> BNT5 -2.06261646 -0.81597484 -0.56826128 0.53560509 -0.7971357244
#> BOF1 1.81480172 -0.09947140 -0.03338089 0.96823582 0.7194896725
#> BOT3 -0.42652849 -0.95308183 -1.45302133 -0.12372612 -0.2651981743
#> CAR1 -0.17585046 -1.87750502 0.22590842 1.03635154 -0.0008125122
#> CTR7 2.13665072 -0.46384719 -0.65813683 1.53673955 1.5430384989
#> CYR26 -2.53285041 0.80770201 -1.35332082 0.24562661 1.2458162395
#> FCT13 1.31708732 2.36303397 0.54897789 -2.15437994 -0.8175669328
#> HAY2 -1.64251639 0.78682069 -0.47047066 -0.83641880 0.6454900542
#> HHR2 1.51967283 0.04742786 0.02391089 1.42066544 2.6872436304
#> HNT18 -1.52995055 0.46901980 0.97973561 1.42193003 -1.3638930576
#> HOP1 1.02775048 -1.79937485 1.48745613 -1.01928580 -2.6544788183
#> INK1 2.65595330 0.46573292 0.49661748 0.49979492 -0.9055914561
#> INK6 2.65595330 0.46573292 0.49661748 0.40724720 -1.3231796846
#> KBG6 -0.88092743 0.88360393 2.43538899 0.37907290 1.5493482879
#> KRV3 -0.86020674 0.60638611 3.28311356 0.95403400 1.3047334526
#> LBH40 -0.39148006 3.58236443 -0.86979703 -0.52049296 0.7182661429
#> LCV19 -2.10472186 -0.72979114 -0.84393462 0.95477464 -1.2752955004
#> LFV10 -0.70247745 -2.15123119 -2.52013644 0.09156243 1.6031745397
#> LTF1 -1.64661629 0.43065330 0.40152506 -1.90495327 0.5977506625
#> LTF6 -1.64661629 0.43065330 0.40152506 -1.58765481 0.2282181833
#> MLG2 0.79400659 -1.68422011 -0.79964612 -0.85647358 0.3750514845
#> MLG5 0.79400659 -1.68422011 -0.79964612 -1.59764781 -0.0074023270
#> MNR2 -1.64833305 -0.70076694 2.16224559 -0.57992342 1.1094446997
#> MRO3 0.84533451 0.32433402 -1.29861325 0.92793262 1.4134687641
#> OLA14 -0.81445084 0.14539414 0.16961964 1.57457009 -1.8896490327
#> OLA9 -0.81445084 0.14539414 0.16961964 1.95997884 -3.1707018189
#> POT1 -0.81296333 -1.84701615 -2.86663451 0.38807069 2.8330896133
#> PRU1 -0.25228743 -0.08106274 1.27406955 0.91229411 -0.4608526034
#> PRU6 -0.25228743 -0.08106274 1.27406955 0.68651209 -0.5116435167
#> PT8 -1.83991210 -2.14713214 2.29618501 -0.12200635 0.9958066956
#> REY5 -0.47238661 -1.30459666 -0.18793822 1.24674474 0.1156194052
#> RPS38 -1.01070710 0.04570288 -0.71027094 2.01997659 -1.6000988948
#> RPS9 -1.01070710 0.04570288 -0.71027094 1.62378263 -2.0160940939
#> SBR10 -0.58648526 1.76832530 -1.11483259 1.33384428 0.0947524518
#> SKY3 -0.31050081 -1.47637523 -0.40355817 -0.39995703 -0.3835032518
#> SKY5 -0.31050081 -1.47637523 -0.40355817 -0.35525520 -0.7312027157
#> SRM1 0.63080315 -0.69734602 0.02683349 -1.61571066 -0.1729069906
#> SRM7 0.63080315 -0.69734602 0.02683349 -2.24534752 -0.0054536726
#> SUT7 -0.34837060 0.75112390 2.02458858 -0.84520937 -1.0269243787
#> TTC1 1.27637074 -0.06280160 -0.14048261 0.93636800 1.0276256196
#> TTC2 1.27637074 -0.06280160 -0.14048261 0.98248003 1.2805980822
#> WAW1 2.04864589 0.07040988 0.12994167 1.24710313 0.6633282699
#> WDC1 0.93464434 0.66493350 1.78723529 0.64218008 0.3561911663
#> WDC2 0.93464434 0.66493350 1.78723529 1.02490718 1.9661662864
#> WHR2 1.00755057 -0.15652483 -0.88109976 -0.89529210 -1.9552230969
#> WTW14 1.30717107 -0.64369150 0.32924405 -1.29521166 -1.2772384337
#> WTW3 1.30717107 -0.64369150 0.32924405 -1.17126810 -0.4047634964
#> YUC3 0.05878549 0.55751783 1.36710582 -2.14525585 1.1662467386
#> ZUM18 -0.08810396 3.02526110 -1.30740035 0.40585515 0.0614906320
#> ZUM20 -0.08810396 3.02526110 -1.30740035 0.39297734 0.0751097215
#> PC3
#> ALT3 1.36393839
#> BAR360 0.32928135
#> BLL5 0.32307359
#> BNT5 0.92230946
#> BOF1 0.00126034
#> BOT3 -1.77036971
#> CAR1 1.28343963
#> CTR7 -0.39837430
#> CYR26 0.45356677
#> FCT13 -1.54412883
#> HAY2 -3.35610236
#> HHR2 -0.48730308
#> HNT18 0.03353609
#> HOP1 3.14963379
#> INK1 -1.57155262
#> INK6 -0.93628485
#> KBG6 0.92436500
#> KRV3 1.43165679
#> LBH40 0.13490647
#> LCV19 0.62060155
#> LFV10 -0.08982798
#> LTF1 -0.49057531
#> LTF6 0.62070618
#> MLG2 0.41294351
#> MLG5 1.65400213
#> MNR2 -1.50388330
#> MRO3 1.19085326
#> OLA14 1.03326291
#> OLA9 -2.22443495
#> POT1 1.00535396
#> PRU1 1.97510046
#> PRU6 0.43246364
#> PT8 -1.77439014
#> REY5 1.21030440
#> RPS38 -0.92152389
#> RPS9 -1.82713565
#> SBR10 1.01613971
#> SKY3 -2.59756340
#> SKY5 -0.73327785
#> SRM1 -1.08994543
#> SRM7 -0.88334213
#> SUT7 1.74380282
#> TTC1 0.18045650
#> TTC2 -0.14274102
#> WAW1 0.64676783
#> WDC1 -0.04577562
#> WDC2 -1.10844126
#> WHR2 1.93142556
#> WTW14 -0.48709556
#> WTW3 -0.86315517
#> YUC3 -0.19366717
#> ZUM18 0.94898317
#> ZUM20 0.06675630
#>
#> $biplot
#> RDA1 RDA2 RDA3 PC1 PC2 PC3
#> CA_rPCA1 -0.2344634 0.4008704 0.82274815 0 0 0
#> CA_rPCA2 0.2021008 0.3349104 -0.08966561 0 0 0
#> CA_rPCA3 0.6824054 -0.2888954 0.51661155 0 0 0
#>
#> $call
#> rda(formula = gen ~ CA_rPCA1 + CA_rPCA2 + CA_rPCA3 + Condition(x +
#> y), data = moddf)
#>
#> $tot.chi
#> [1] 143.389Run a partial RDA with population genetic structure and geography as covariables
Lastly, let’s run a partial RDA with both geography
(correctGEO = TRUE) and population structure
(correctPC = TRUE) as conditioning variables. Within
algatr, population structure is quantified using a PCA, and a user can
specify the number of axes to retain in the RDA model. In this case,
we’ll retain three axes, which we can specify using the nPC
argument. If this argument is set to "manual", a screeplot
will be printed and a user can manually enter the number of PCs that
they feel best describe the data.
mod_pRDA_gs <- rda_run(gen, env, liz_coords,
model = "full",
correctGEO = TRUE,
correctPC = TRUE,
nPC = 3
)
Now, we see that even more inertia/variance is explained by the conditioning variables.
head(summary(mod_pRDA_gs))
#> $species
#> RDA1 RDA2 RDA3 PC1
#> Locus_10 -1.265110e-02 -1.061950e-02 2.431077e-02 9.692674e-03
#> Locus_15 7.156365e-02 -4.413745e-02 -7.565983e-02 1.528115e-01
#> Locus_22 6.219065e-03 1.820773e-04 1.421059e-02 3.812177e-02
#> Locus_28 7.276209e-02 6.050645e-02 -7.430354e-02 -2.306247e-02
#> Locus_32 -1.465809e-02 1.498110e-03 2.902435e-02 -1.417281e-02
#> Locus_35 5.442256e-32 -2.051344e-33 -4.397205e-32 8.447652e-17
#> Locus_37 -1.471592e-32 2.665310e-32 -3.475241e-32 2.963656e-17
#> Locus_61 5.079288e-02 -8.662815e-02 1.033709e-02 -1.499091e-01
#> Locus_71 -3.549646e-03 7.653060e-03 7.707053e-03 1.940609e-02
#> Locus_72 -1.773829e-32 -5.166490e-32 3.772338e-32 -7.726959e-17
#> Locus_82 -4.980745e-32 -2.499226e-32 -1.121216e-32 4.944460e-17
#> Locus_84 -3.418642e-02 1.226410e-02 -1.322891e-02 1.917286e-03
#> Locus_90 -7.282587e-02 -1.774081e-02 9.560021e-02 -6.972555e-02
#> Locus_91 -4.579609e-03 -2.485802e-02 3.502555e-03 2.345279e-02
#> Locus_96 1.892074e-32 -4.702379e-33 -4.023105e-32 -1.269522e-17
#> Locus_104 -5.749339e-04 2.282730e-02 1.711713e-03 -9.402803e-03
#> Locus_122 8.296874e-02 -5.828088e-02 -1.623578e-02 -2.118480e-02
#> Locus_125 -2.582196e-02 -1.053880e-01 4.443282e-02 5.487456e-02
#> Locus_131 -6.857927e-02 1.165864e-02 3.259892e-03 1.617366e-02
#> Locus_133 -1.324420e-34 8.146636e-33 1.419107e-33 -1.343930e-17
#> Locus_138 1.520506e-02 -9.886540e-02 4.227453e-02 -1.411905e-01
#> Locus_154 5.312751e-02 -1.710234e-02 4.132451e-02 1.120578e-01
#> Locus_162 5.283444e-02 -9.120378e-02 6.621954e-02 8.621438e-02
#> Locus_166 -2.143177e-02 -5.762018e-02 4.069952e-02 1.119030e-02
#> Locus_177 3.170033e-33 6.781091e-33 -3.127576e-33 -3.306783e-17
#> Locus_185 7.661909e-33 -4.428586e-33 3.805518e-33 -3.887761e-17
#> Locus_191 -1.286253e-01 -7.085994e-03 -5.372690e-02 -3.871974e-02
#> Locus_193 -3.006776e-02 4.452191e-02 -2.604686e-02 4.377229e-02
#> Locus_207 1.264087e-02 -1.010820e-01 -2.619588e-02 -7.643039e-02
#> Locus_211 -1.718575e-03 -1.228035e-03 -1.832266e-05 6.572077e-03
#> Locus_213 1.243906e-02 5.993000e-02 -7.456160e-02 -4.193926e-02
#> Locus_214 5.875045e-02 -8.860422e-03 -7.407401e-02 1.266364e-02
#> Locus_222 -3.888618e-02 1.713755e-02 -1.218347e-03 -6.741545e-03
#> Locus_227 -1.222001e-02 -9.826129e-02 3.674431e-02 1.960009e-01
#> Locus_242 -1.945394e-02 -2.723370e-02 -1.132344e-03 -5.808013e-02
#> Locus_249 1.290936e-02 -9.534522e-02 7.434976e-03 -1.873046e-01
#> Locus_254 3.256440e-02 2.318649e-02 1.060373e-02 -5.516323e-02
#> Locus_256 6.844595e-02 -7.164116e-02 1.072068e-01 -1.656455e-01
#> Locus_263 -3.524166e-02 -5.667914e-02 -4.499779e-02 -1.041301e-01
#> Locus_268 3.163515e-02 -2.294814e-02 -1.997833e-02 -1.632825e-02
#> Locus_279 1.969771e-02 4.674535e-02 -5.420942e-02 -1.846826e-02
#> Locus_285 -7.993239e-02 -3.400796e-02 3.858166e-02 -2.818454e-02
#> Locus_287 -7.318880e-03 -3.607567e-02 -1.959786e-02 4.442494e-02
#> Locus_292 5.283150e-02 3.989373e-02 -7.620603e-03 -2.441333e-02
#> Locus_294 -3.018238e-02 3.273629e-03 1.646057e-02 9.476250e-04
#> Locus_303 -7.461399e-34 -3.578388e-35 -9.765874e-34 1.216400e-29
#> Locus_308 2.848947e-02 5.885215e-02 4.453450e-02 2.855331e-02
#> Locus_312 -1.003019e-01 -6.467156e-03 -5.669139e-02 -4.051785e-02
#> Locus_313 -7.542955e-34 -3.701299e-34 -9.655292e-34 1.537570e-31
#> Locus_314 -2.905995e-02 6.866070e-03 1.864342e-02 3.695970e-02
#> Locus_317 -2.518729e-02 2.663515e-02 1.184095e-02 -2.767328e-04
#> Locus_318 1.042608e-01 -5.125456e-02 7.515167e-02 -4.483574e-02
#> Locus_323 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_330 2.443710e-02 -1.821699e-03 -7.522013e-03 3.766558e-03
#> Locus_331 -5.004530e-03 2.896309e-03 -2.638515e-02 -1.440485e-02
#> Locus_333 1.682096e-02 1.631427e-02 1.437705e-02 -6.271966e-03
#> Locus_334 1.682150e-03 -1.840840e-02 9.729215e-03 -1.258725e-02
#> Locus_336 8.276465e-04 -1.373907e-02 6.658967e-03 2.527783e-02
#> Locus_337 -2.300557e-02 -5.569232e-02 -8.420855e-03 -1.597356e-02
#> Locus_339 8.719110e-02 -1.155054e-01 -7.705311e-02 8.625817e-03
#> Locus_341 5.126243e-03 -1.303852e-01 -4.282405e-02 4.469000e-02
#> Locus_345 -5.665904e-02 2.573889e-03 -3.515449e-02 1.037990e-02
#> Locus_350 -3.927393e-02 -1.103370e-01 2.877912e-02 -4.013863e-02
#> Locus_354 -2.798177e-02 -1.840401e-02 -3.769987e-02 1.157464e-02
#> Locus_359 1.197842e-01 3.348661e-02 -1.274999e-02 1.845968e-01
#> Locus_365 1.465694e-02 -8.096988e-03 -5.233872e-03 3.046243e-02
#> Locus_366 4.570012e-02 -2.007474e-02 1.930442e-03 1.580313e-02
#> Locus_369 2.195991e-02 1.349144e-02 -1.991360e-02 -2.302162e-02
#> Locus_372 -1.300721e-02 3.349400e-02 -3.374034e-03 -1.089340e-03
#> Locus_389 7.380489e-02 4.136246e-02 4.783394e-03 2.238237e-02
#> Locus_391 -1.546256e-03 -1.246929e-02 -5.940525e-03 1.616547e-02
#> Locus_392 1.401800e-02 2.813915e-04 -2.143733e-02 2.531885e-02
#> Locus_393 1.107515e-02 1.477904e-02 5.541549e-03 -9.800844e-03
#> Locus_394 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_396 -1.812891e-02 -3.205908e-02 2.165487e-02 4.907840e-02
#> Locus_399 -4.046509e-04 -3.512021e-02 -1.269367e-02 2.929239e-02
#> Locus_400 -4.509222e-02 -1.174961e-02 -3.868027e-02 1.134709e-03
#> Locus_401 -2.483750e-02 -1.756725e-02 1.042686e-01 4.623969e-02
#> Locus_404 5.758560e-02 1.739663e-02 -4.555488e-02 2.066394e-01
#> Locus_406 -5.867468e-02 7.799126e-02 2.199701e-02 3.282976e-02
#> Locus_409 2.543492e-02 2.150112e-02 -2.385539e-02 3.408232e-02
#> Locus_420 2.128878e-02 -2.726923e-02 2.594354e-02 1.207372e-02
#> Locus_422 1.487984e-02 3.174091e-02 1.791766e-02 -4.190031e-03
#> Locus_427 8.122980e-02 -3.229559e-02 -2.865649e-02 1.171061e-01
#> Locus_430 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_436 -5.268985e-02 4.190204e-02 1.346555e-02 -2.265667e-03
#> Locus_438 5.040780e-02 -1.886911e-02 -2.712564e-02 -4.598600e-02
#> Locus_439 7.576080e-02 2.063751e-02 5.540478e-02 2.742758e-02
#> Locus_441 -4.750655e-02 1.905673e-02 3.728687e-03 -1.587248e-01
#> Locus_445 -2.153312e-02 2.762316e-02 1.071359e-02 -2.664342e-04
#> Locus_448 -2.594165e-02 8.837511e-03 1.052179e-02 7.995941e-03
#> Locus_449 2.258996e-02 1.956144e-03 -1.883625e-02 9.932985e-02
#> Locus_456 -4.412527e-02 9.628132e-03 -3.342726e-02 -2.813406e-02
#> Locus_457 -8.075424e-02 2.073774e-02 -1.774694e-02 2.231610e-03
#> Locus_458 1.018585e-02 -6.812193e-02 8.577398e-03 -2.100859e-02
#> Locus_460 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_461 -8.248044e-03 3.792477e-02 5.076561e-04 2.210900e-02
#> Locus_462 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_464 3.982177e-02 4.233318e-02 2.263375e-02 1.407842e-02
#> Locus_469 2.006281e-01 -3.066340e-03 3.506400e-02 3.772639e-02
#> Locus_475 -3.682552e-02 -9.547344e-03 -3.398619e-02 -5.112033e-02
#> Locus_480 1.964524e-02 2.551410e-02 -6.478240e-02 3.824114e-02
#> Locus_482 3.084929e-02 -6.063062e-03 1.174640e-02 -7.069478e-02
#> Locus_483 -2.400874e-02 -1.941891e-02 -3.256460e-02 -8.662728e-03
#> Locus_491 2.770903e-02 2.378914e-02 -1.611962e-02 -3.006028e-02
#> Locus_498 2.542042e-03 2.647375e-02 1.476851e-02 8.464816e-03
#> Locus_500 1.597473e-02 -7.542226e-04 -7.481974e-03 -1.027680e-02
#> Locus_503 -2.722103e-02 2.957204e-02 -1.819108e-02 1.053492e-01
#> Locus_505 3.562588e-02 7.127157e-02 2.764397e-02 1.240265e-01
#> Locus_508 4.229096e-02 -5.053905e-02 -3.909680e-02 2.106382e-01
#> Locus_512 -1.881951e-02 -7.987730e-02 3.373229e-02 3.586261e-02
#> Locus_516 -1.105337e-02 -1.174118e-02 -1.632869e-02 -8.375382e-03
#> Locus_517 -9.244477e-02 4.870668e-03 -4.568534e-03 -4.597650e-02
#> Locus_518 1.714581e-02 1.569782e-03 -6.349146e-04 1.445441e-02
#> Locus_528 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_529 3.422287e-02 -4.721163e-02 3.816820e-02 -7.594499e-02
#> Locus_531 6.982119e-03 4.772175e-02 3.654254e-03 2.457008e-02
#> Locus_532 7.569738e-03 4.078215e-02 4.262411e-02 2.776842e-02
#> Locus_533 7.723959e-03 -6.127555e-03 -1.672778e-02 2.762727e-04
#> Locus_535 -1.732322e-02 3.041475e-02 2.596224e-02 2.502921e-02
#> Locus_546 3.917409e-02 7.985946e-02 3.487116e-02 7.722801e-02
#> Locus_557 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_558 6.963439e-03 -4.571689e-02 5.093808e-02 -2.720982e-02
#> Locus_561 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_562 -3.136185e-02 4.385659e-02 -1.277833e-02 -4.473841e-04
#> Locus_564 -2.450783e-02 1.244893e-02 1.670893e-02 2.854246e-02
#> Locus_566 -7.641597e-02 1.071791e-02 -1.156890e-02 -2.075618e-02
#> Locus_572 -2.410767e-02 -1.351173e-02 -3.971735e-02 3.467179e-02
#> Locus_573 -2.666289e-02 2.066910e-03 -1.474387e-02 -2.099082e-03
#> Locus_579 3.026130e-02 1.266862e-02 -2.341864e-02 3.539560e-02
#> Locus_581 -7.374384e-02 -1.687129e-02 -4.780024e-02 -2.162289e-02
#> Locus_582 -2.666289e-02 2.066910e-03 -1.474387e-02 -2.099082e-03
#> Locus_583 2.244614e-02 9.782317e-02 1.989585e-02 -3.580367e-02
#> Locus_586 -4.288935e-02 -8.115918e-03 9.953219e-03 -2.867889e-02
#> Locus_589 -3.597188e-02 -6.021802e-02 -8.174159e-03 -1.180252e-01
#> Locus_596 4.284556e-02 -9.763801e-03 -1.054933e-02 1.536697e-02
#> Locus_597 -5.360240e-03 -6.566014e-02 5.220959e-03 3.996084e-02
#> Locus_598 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_600 1.451375e-02 -5.535598e-03 2.642291e-02 -1.814687e-03
#> Locus_602 -1.786434e-02 6.353564e-02 2.163798e-02 -6.108976e-02
#> Locus_603 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_604 2.083298e-02 -4.510854e-03 3.086302e-03 -1.358493e-02
#> Locus_609 5.672780e-02 1.005084e-01 -1.796862e-02 -8.485347e-02
#> Locus_612 2.482526e-02 -2.298542e-02 -1.653594e-02 -9.854653e-02
#> Locus_613 -2.334008e-02 2.497179e-02 5.409478e-02 -2.454654e-02
#> Locus_614 1.342500e-02 -3.900458e-03 -1.507407e-03 1.045998e-02
#> Locus_622 1.672908e-02 1.704342e-02 -6.162813e-03 -1.854947e-02
#> Locus_625 3.661046e-03 -1.117891e-03 3.136776e-02 -2.175391e-02
#> Locus_626 5.541807e-02 4.757827e-02 -3.223923e-02 -6.012056e-02
#> Locus_633 1.721654e-02 -9.057445e-03 -9.821835e-03 -1.785460e-02
#> Locus_634 1.711144e-02 -2.360582e-02 1.908410e-02 -3.797249e-02
#> Locus_635 -6.123672e-02 7.665292e-02 3.426424e-03 1.464017e-02
#> Locus_636 -5.830641e-02 -1.734899e-01 -7.327541e-02 9.308276e-03
#> Locus_637 1.102643e-02 -5.307076e-02 9.674018e-03 -5.773936e-02
#> Locus_639 5.011602e-02 -1.194662e-02 -1.738389e-02 -4.468935e-05
#> Locus_642 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_647 -2.468729e-03 -8.515756e-03 1.357176e-02 6.161016e-02
#> Locus_652 2.787570e-02 -2.213634e-02 1.335934e-02 1.175272e-02
#> Locus_665 1.834323e-04 -5.291478e-03 -1.110999e-04 -1.515960e-03
#> Locus_671 -8.125766e-03 -4.640478e-02 -2.565375e-02 6.527572e-02
#> Locus_679 2.111377e-02 -1.939056e-02 -2.304468e-02 5.357261e-02
#> Locus_681 4.390094e-02 -5.034683e-02 -3.246038e-04 -3.014023e-02
#> Locus_682 -8.125766e-03 -4.640478e-02 -2.565375e-02 6.527572e-02
#> Locus_683 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_685 -2.666289e-02 2.066910e-03 -1.474387e-02 -2.099082e-03
#> Locus_687 -2.484107e-02 -3.216837e-03 -8.853759e-03 1.293327e-02
#> Locus_691 -6.416881e-02 8.526775e-03 5.045300e-03 9.906250e-03
#> Locus_699 9.562269e-02 7.522103e-02 -1.157062e-01 -1.504315e-02
#> Locus_701 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_707 3.566324e-02 1.125402e-02 -2.561567e-02 -1.211277e-02
#> Locus_710 1.273844e-04 -2.536210e-03 -6.980962e-02 -2.725225e-02
#> Locus_711 6.129784e-03 2.930805e-02 2.816793e-02 -3.192525e-02
#> Locus_712 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_717 2.111377e-02 -1.939056e-02 -2.304468e-02 5.357261e-02
#> Locus_718 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_724 2.102862e-02 -1.091094e-01 3.006196e-02 3.120779e-02
#> Locus_725 1.711144e-02 -2.360582e-02 1.908410e-02 -3.797249e-02
#> Locus_727 5.391836e-04 -2.182421e-02 -6.144403e-03 6.561478e-03
#> Locus_729 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_733 2.478753e-02 1.399340e-02 5.750521e-03 -7.708163e-02
#> Locus_734 1.716665e-02 3.611204e-04 -6.874523e-04 1.917822e-02
#> Locus_735 2.180319e-02 -6.917378e-03 5.587083e-02 -1.287338e-01
#> Locus_736 -8.125766e-03 -4.640478e-02 -2.565375e-02 6.527572e-02
#> Locus_740 1.567767e-01 9.050585e-02 -9.366153e-02 1.103964e-01
#> Locus_742 -4.157559e-02 1.990819e-02 -2.311843e-02 4.153107e-02
#> Locus_746 -2.078663e-02 2.474601e-02 8.600397e-02 2.857834e-02
#> Locus_750 -4.124022e-03 1.896239e-02 2.538280e-04 1.105450e-02
#> Locus_751 3.345816e-02 3.408683e-02 -1.232563e-02 -3.709894e-02
#> Locus_753 -4.901902e-02 -4.997920e-03 -3.425330e-02 -1.204677e-02
#> Locus_758 -6.085002e-03 -2.946494e-02 -9.410080e-03 -1.976686e-02
#> Locus_765 -1.185234e-02 1.017243e-02 -3.458657e-03 -4.011668e-02
#> Locus_769 -1.369562e-02 1.455224e-02 4.585299e-02 -5.142365e-02
#> Locus_770 1.404629e-01 2.469576e-03 -5.726887e-02 7.241997e-02
#> Locus_771 7.495368e-02 -2.251469e-02 -9.565812e-02 6.511456e-02
#> Locus_776 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_779 -1.082224e-01 2.124108e-02 -3.583965e-02 6.431389e-03
#> Locus_784 -1.127248e-02 5.680763e-03 -2.291526e-02 -1.987730e-02
#> Locus_786 3.286568e-02 4.746215e-02 -9.012516e-03 -3.343566e-02
#> Locus_787 2.806479e-02 7.256401e-03 -2.215292e-02 -2.198284e-02
#> Locus_798 -3.649254e-02 4.323386e-02 -4.122541e-02 1.127620e-01
#> Locus_802 5.587447e-04 -4.162943e-02 4.030008e-02 1.452597e-01
#> Locus_803 -9.551458e-03 -2.153891e-02 4.340230e-03 -4.007158e-02
#> Locus_807 -8.248044e-03 3.792477e-02 5.076561e-04 2.210900e-02
#> Locus_808 1.452297e-03 -8.397265e-03 -4.419596e-02 8.705322e-03
#> Locus_809 3.345816e-02 3.408683e-02 -1.232563e-02 -3.709894e-02
#> Locus_811 4.935799e-02 1.348748e-03 -1.625878e-02 -1.028388e-02
#> Locus_816 -3.251702e-02 2.515851e-03 8.553248e-04 2.036854e-03
#> Locus_821 8.462378e-03 -1.067476e-03 -4.003902e-05 1.404336e-02
#> Locus_825 6.489305e-02 -4.658467e-02 -3.827005e-02 1.008501e-01
#> Locus_831 -3.606081e-02 8.564627e-02 -1.081279e-02 1.204314e-03
#> Locus_835 2.290374e-02 -3.864087e-02 4.357646e-02 -3.437094e-02
#> Locus_838 4.261299e-03 -4.196882e-02 -1.240378e-02 3.268009e-02
#> Locus_847 -6.179793e-02 7.160482e-02 -2.227751e-02 -7.733220e-05
#> Locus_850 -6.563856e-02 3.966337e-03 -1.287850e-02 -1.552423e-02
#> Locus_851 -1.190504e-03 -2.900203e-02 -1.197383e-02 2.424291e-02
#> Locus_853 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_854 1.781294e-02 3.563578e-02 1.382199e-02 6.201326e-02
#> Locus_859 -2.582196e-02 -1.053880e-01 4.443282e-02 5.487456e-02
#> Locus_861 -1.036899e-01 5.120426e-03 -4.374170e-02 -9.653317e-04
#> Locus_864 5.727560e-03 4.904979e-03 1.988956e-02 6.193910e-02
#> Locus_869 8.484063e-04 -3.780726e-02 7.262042e-03 7.140643e-02
#> Locus_870 -5.744981e-02 2.309621e-02 -2.923391e-02 6.856337e-03
#> Locus_875 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_876 -4.011073e-02 3.836715e-02 -3.148089e-02 7.471089e-02
#> Locus_879 6.948743e-02 6.911193e-03 -1.862393e-02 -1.150690e-02
#> Locus_881 7.266934e-02 3.240528e-02 3.982847e-02 9.705821e-02
#> Locus_887 -1.155458e-02 -1.649627e-02 3.911193e-02 4.055481e-03
#> Locus_888 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_895 2.908705e-02 -2.549269e-04 -3.680149e-02 5.827429e-03
#> Locus_896 7.297299e-03 1.263692e-03 -6.823658e-03 3.835973e-02
#> Locus_897 -2.055062e-02 -2.514710e-02 -2.905351e-03 1.578032e-02
#> Locus_898 -1.513092e-02 4.184698e-03 -2.033321e-02 -6.387923e-02
#> Locus_899 2.573113e-02 -8.404072e-02 4.354381e-02 1.628640e-02
#> Locus_901 6.905980e-02 -9.122548e-02 -4.747665e-02 7.757238e-02
#> Locus_902 3.612950e-02 -5.282104e-02 9.694352e-04 2.866027e-02
#> Locus_903 -3.963687e-02 5.873199e-02 -1.541192e-02 6.886960e-02
#> Locus_905 3.905253e-02 -9.390524e-04 -3.753116e-03 -3.323095e-02
#> Locus_908 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_909 3.665710e-03 -2.249774e-02 2.970623e-02 1.399653e-02
#> Locus_915 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_924 -6.761655e-02 -5.711273e-02 -4.069412e-02 -1.455894e-02
#> Locus_932 -1.472332e-02 3.122702e-02 3.458457e-02 -2.321277e-03
#> Locus_935 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_936 7.756319e-02 9.382347e-02 8.891549e-02 7.576986e-02
#> Locus_938 3.422287e-02 -4.721163e-02 3.816820e-02 -7.594499e-02
#> Locus_941 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_944 1.355213e-02 4.441983e-02 -1.208181e-02 -2.087521e-02
#> Locus_945 3.092601e-02 7.473186e-03 -3.942663e-02 2.585602e-02
#> Locus_946 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_949 4.888930e-02 2.394065e-03 1.760904e-02 4.908530e-03
#> Locus_950 1.207734e-02 5.696972e-03 -2.561539e-02 1.755020e-02
#> Locus_951 1.868630e-02 7.228949e-02 -1.810242e-02 3.325818e-01
#> Locus_953 6.907326e-02 2.567310e-02 -2.209731e-02 2.866206e-02
#> Locus_958 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_965 3.482298e-02 3.509850e-02 -2.370674e-02 -1.319128e-02
#> Locus_966 -9.698966e-04 -2.081986e-02 -6.411547e-03 1.717425e-02
#> Locus_967 2.682929e-02 2.034125e-02 -3.067541e-02 5.838027e-03
#> Locus_968 -1.622723e-02 3.710568e-02 4.700054e-02 -3.409407e-02
#> Locus_970 -3.975034e-02 -5.290877e-02 -1.345343e-02 -2.854000e-03
#> Locus_972 2.663142e-02 -1.266939e-02 6.619614e-03 2.694139e-02
#> Locus_974 -3.078692e-02 2.102930e-02 -1.449004e-02 8.955419e-03
#> Locus_979 2.312288e-02 -1.246210e-02 -1.080391e-02 6.661210e-03
#> Locus_992 2.848358e-02 -1.101730e-02 1.175228e-02 3.365773e-02
#> Locus_998 7.322092e-03 -2.235782e-03 6.273552e-02 -4.350783e-02
#> Locus_999 1.262653e-02 1.752213e-02 6.733573e-02 1.958632e-02
#> Locus_1002 -2.833744e-02 -3.052990e-02 -2.236854e-02 -2.813335e-02
#> Locus_1005 6.992198e-02 -5.666709e-03 -8.157030e-02 7.406262e-02
#> Locus_1006 -1.991921e-02 3.281350e-02 2.383279e-02 4.494784e-02
#> Locus_1007 -1.772778e-02 -6.844674e-02 6.759909e-02 6.719287e-02
#> Locus_1015 -1.047901e-02 1.948296e-02 2.054241e-02 2.966784e-02
#> Locus_1016 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1017 -7.934193e-03 4.848218e-03 -4.202367e-03 5.956739e-02
#> Locus_1023 1.711144e-02 -2.360582e-02 1.908410e-02 -3.797249e-02
#> Locus_1024 -5.710411e-02 6.931152e-04 -2.770838e-02 -5.240405e-03
#> Locus_1025 -1.254562e-02 3.552149e-02 2.729240e-02 -5.222619e-02
#> Locus_1027 -7.002446e-03 -2.551073e-02 1.070052e-02 1.901195e-02
#> Locus_1030 9.303365e-02 -7.407730e-03 -7.814644e-02 6.298710e-02
#> Locus_1031 -5.848715e-03 -2.451178e-02 -6.585570e-02 -3.105134e-02
#> Locus_1033 3.175864e-03 -2.160205e-02 2.154127e-02 1.217085e-02
#> Locus_1034 -5.125148e-03 5.780690e-02 -5.026300e-02 2.354077e-02
#> Locus_1035 5.763591e-04 -8.350568e-03 -4.710218e-04 1.008781e-03
#> Locus_1038 2.083298e-02 -4.510854e-03 3.086302e-03 -1.358493e-02
#> Locus_1041 -5.171553e-02 4.065827e-02 4.405081e-02 1.922464e-02
#> Locus_1042 -1.280905e-02 2.749139e-02 -3.540429e-02 -1.745577e-02
#> Locus_1044 5.921606e-02 7.190078e-02 2.273486e-02 1.367869e-01
#> Locus_1046 -5.908534e-03 -2.718328e-02 1.151589e-02 8.425332e-03
#> Locus_1051 -1.291098e-02 -5.269402e-02 2.221641e-02 2.743728e-02
#> Locus_1054 2.243155e-02 -1.204780e-02 -2.529279e-02 1.283338e-02
#> Locus_1055 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1065 -1.453071e-02 -1.038100e-03 -4.078527e-02 -4.142507e-02
#> Locus_1071 -1.217000e-02 -5.892989e-02 -1.882016e-02 -3.953373e-02
#> Locus_1072 -1.082844e-01 1.019388e-02 6.240143e-03 9.546950e-03
#> Locus_1073 3.811406e-02 5.336215e-02 -2.436470e-02 -5.097390e-02
#> Locus_1074 -3.204654e-05 -3.302519e-02 -4.374802e-03 1.171580e-02
#> Locus_1075 2.154421e-02 1.504089e-02 1.581047e-02 3.056497e-02
#> Locus_1076 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1080 1.210691e-02 -2.070951e-02 -7.301053e-03 -5.237735e-02
#> Locus_1082 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1093 5.787026e-02 -9.137294e-03 -2.543320e-02 -3.327334e-02
#> Locus_1095 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1098 -1.369562e-02 1.455224e-02 4.585299e-02 -5.142365e-02
#> Locus_1099 8.276465e-04 -1.373907e-02 6.658967e-03 2.527783e-02
#> Locus_1105 -6.109647e-02 -6.610917e-02 7.440798e-02 -1.272299e-01
#> Locus_1106 -5.387065e-02 4.089784e-03 -2.624698e-03 8.293237e-03
#> Locus_1107 -3.741650e-03 -4.261579e-03 -8.199546e-04 -8.718234e-03
#> Locus_1108 -5.332579e-02 4.133821e-03 -2.948774e-02 -4.198165e-03
#> Locus_1109 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1113 7.441619e-03 1.094048e-02 1.744985e-03 7.988862e-03
#> Locus_1115 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1118 -2.655773e-03 -2.551214e-02 1.037990e-02 1.155304e-02
#> Locus_1122 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1127 -2.666289e-02 2.066910e-03 -1.474387e-02 -2.099082e-03
#> Locus_1138 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1141 -6.759634e-02 4.151336e-02 1.807697e-02 -6.502462e-02
#> Locus_1143 3.468812e-02 2.804361e-02 -7.016501e-02 -8.625993e-03
#> Locus_1144 7.173268e-02 -4.696376e-02 -3.999670e-02 -1.861315e-02
#> Locus_1155 -3.750355e-03 -2.870717e-02 6.195162e-03 -2.085609e-02
#> Locus_1158 -1.109895e-01 -7.609115e-03 -4.695979e-02 -2.797181e-02
#> Locus_1159 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1161 -1.535142e-03 -6.519513e-03 -1.294225e-04 5.056117e-03
#> Locus_1162 4.292198e-02 2.942300e-02 -8.024500e-02 -4.837501e-03
#> Locus_1166 -3.877978e-04 -1.649245e-02 1.658500e-03 3.638360e-03
#> Locus_1172 7.974724e-02 -3.029778e-02 -1.078296e-02 8.658953e-02
#> Locus_1179 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1181 -5.908534e-03 -2.718328e-02 1.151589e-02 8.425332e-03
#> Locus_1186 1.959837e-02 -9.063269e-02 -4.969934e-03 -4.704379e-02
#> Locus_1191 2.334647e-03 7.577779e-04 1.560524e-02 -1.089229e-03
#> Locus_1197 2.215030e-02 2.955809e-02 1.108310e-02 -1.960169e-02
#> Locus_1199 1.586368e-02 -1.341580e-02 2.110644e-02 1.711545e-02
#> Locus_1201 5.331656e-02 1.500478e-02 -4.063589e-02 2.075635e-01
#> Locus_1204 3.163515e-02 -2.294814e-02 -1.997833e-02 -1.632825e-02
#> Locus_1206 -1.127248e-02 5.680763e-03 -2.291526e-02 -1.987730e-02
#> Locus_1211 -5.975201e-02 3.677485e-02 -2.308997e-02 3.294319e-02
#> Locus_1212 -6.260834e-03 -1.929081e-02 -9.003951e-03 -2.336282e-02
#> Locus_1217 -1.670254e-02 2.661237e-02 6.267410e-02 -5.083118e-02
#> Locus_1223 -2.594165e-02 8.837511e-03 1.052179e-02 7.995941e-03
#> Locus_1224 -1.367744e-02 -1.415466e-02 3.105738e-02 3.250371e-02
#> Locus_1227 -8.776890e-02 8.714171e-03 -1.097570e-02 -3.701615e-02
#> Locus_1231 -5.332579e-02 4.133821e-03 -2.948774e-02 -4.198165e-03
#> Locus_1235 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1236 2.941861e-02 -5.009595e-02 -6.386796e-02 1.257833e-01
#> Locus_1239 2.544362e-02 -6.414146e-02 6.755959e-02 2.182026e-01
#> Locus_1247 1.099537e-01 4.815730e-02 -6.264712e-02 -1.633032e-01
#> Locus_1248 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1249 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1250 -3.070751e-02 1.523827e-02 -1.174905e-02 1.482021e-02
#> Locus_1254 -3.078692e-02 2.102930e-02 -1.449004e-02 8.955419e-03
#> Locus_1255 1.230898e-02 -7.687694e-02 9.361928e-02 -2.772227e-02
#> Locus_1256 -4.309614e-02 -2.660810e-02 1.296516e-02 -6.833678e-02
#> Locus_1258 -3.539694e-02 -8.304531e-02 -9.885873e-03 -1.086224e-01
#> Locus_1262 1.405004e-02 3.330658e-02 -1.706253e-02 1.360305e-02
#> Locus_1263 -6.648067e-02 4.743897e-02 5.607630e-02 -4.366366e-02
#> Locus_1265 1.481769e-02 2.392436e-05 2.561937e-02 9.137820e-03
#> Locus_1266 3.454202e-02 5.267920e-02 7.659174e-03 4.346379e-02
#> Locus_1270 2.070156e-02 1.813250e-02 4.456438e-03 3.738683e-02
#> Locus_1278 3.421849e-02 -2.910704e-02 -1.940708e-02 -5.077555e-03
#> Locus_1279 2.346275e-02 -3.432792e-03 -2.214666e-02 -2.724956e-02
#> Locus_1284 -1.344661e-02 -1.874186e-02 1.084311e-02 1.551321e-02
#> Locus_1286 -2.780856e-02 2.217849e-02 -4.792242e-02 -4.677228e-02
#> Locus_1287 8.303061e-02 3.074969e-02 4.208719e-02 1.327519e-01
#> Locus_1290 1.711144e-02 -2.360582e-02 1.908410e-02 -3.797249e-02
#> Locus_1295 9.706661e-03 -1.053443e-02 6.699692e-03 -1.145320e-03
#> Locus_1301 -1.789999e-02 5.382150e-02 -4.756041e-02 -1.142523e-02
#> Locus_1302 3.110270e-02 4.815964e-04 -1.900741e-02 6.035489e-02
#> Locus_1305 -4.501284e-02 -4.918118e-03 1.350297e-02 2.368808e-02
#> Locus_1313 -4.289069e-02 1.083550e-02 -1.258150e-02 -3.217574e-03
#> Locus_1315 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1316 -4.238597e-02 5.009069e-02 -2.097196e-02 3.828003e-02
#> Locus_1318 3.504614e-02 -4.284611e-02 -1.274811e-02 2.020027e-02
#> Locus_1319 -6.157383e-02 4.205860e-02 -2.898008e-02 1.791084e-02
#> Locus_1338 1.915421e-03 5.294031e-03 8.287313e-03 1.226700e-01
#> Locus_1343 -4.163827e-02 3.928835e-02 3.131065e-02 2.956553e-02
#> Locus_1345 5.292069e-02 -1.283899e-02 4.230264e-03 3.742429e-02
#> Locus_1347 1.953753e-02 1.371157e-02 5.501510e-03 4.242512e-03
#> Locus_1353 -6.384888e-02 1.563137e-02 -3.579620e-03 5.918795e-03
#> Locus_1354 -2.278026e-02 -1.699619e-02 2.870545e-02 -1.391330e-02
#> Locus_1355 3.162294e-03 -1.298129e-02 2.226421e-02 2.418860e-02
#> Locus_1360 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1363 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1364 3.314994e-02 1.964967e-03 6.287991e-03 -3.096358e-02
#> Locus_1366 -7.516844e-02 1.019079e-02 -1.776758e-02 -1.125299e-02
#> Locus_1370 1.179745e-01 -6.225902e-02 -4.104584e-02 6.432303e-02
#> Locus_1374 -1.406087e-02 -5.176174e-02 1.993734e-02 8.274116e-02
#> Locus_1376 -5.522779e-02 8.197264e-03 -2.939496e-02 3.889871e-03
#> Locus_1377 2.921811e-02 -1.764555e-02 -1.022962e-02 3.886995e-02
#> Locus_1379 -1.118487e-01 -1.114539e-02 -6.961228e-02 -1.446766e-02
#> Locus_1380 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1382 -3.146015e-02 -6.603184e-04 3.041461e-02 -7.210361e-02
#> Locus_1384 -5.267225e-02 9.995385e-03 4.756600e-02 3.913219e-03
#> Locus_1392 -4.244004e-02 -6.083276e-02 -5.214179e-03 7.085250e-03
#> Locus_1396 1.063429e-01 9.134259e-02 2.608911e-02 2.597250e-01
#> Locus_1399 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1404 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1408 2.812836e-02 8.227394e-03 4.515893e-02 3.764901e-02
#> Locus_1414 -1.906378e-03 -7.525958e-02 2.319043e-02 -1.032180e-01
#> Locus_1416 5.096541e-02 3.722396e-02 -6.792043e-02 -1.275150e-02
#> Locus_1420 1.565344e-02 -1.090625e-02 -5.874988e-03 5.536265e-02
#> Locus_1423 -2.720434e-02 5.445870e-02 1.542346e-03 8.914751e-02
#> Locus_1424 5.211484e-03 3.742737e-02 -3.850622e-02 1.138580e-01
#> Locus_1425 5.321247e-02 -1.976149e-02 -5.511493e-03 -8.517028e-03
#> Locus_1426 4.281277e-03 -1.405192e-02 -1.762867e-02 -7.224415e-02
#> Locus_1427 1.672908e-02 1.704342e-02 -6.162813e-03 -1.854947e-02
#> Locus_1430 -5.340345e-03 1.728859e-02 -2.698836e-02 1.224230e-01
#> Locus_1433 7.926321e-03 7.920354e-02 4.632642e-02 9.051153e-02
#> Locus_1439 -1.151028e-01 -3.375683e-02 -5.152580e-02 3.819106e-02
#> Locus_1447 -1.088333e-02 -3.060608e-02 8.161413e-02 1.040279e-01
#> Locus_1452 -1.084053e-02 -3.491211e-02 2.596558e-02 7.220688e-02
#> Locus_1455 4.137261e-02 -1.189437e-02 6.969819e-02 9.676089e-02
#> Locus_1457 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1459 -2.781276e-02 4.772150e-02 -1.132044e-02 -2.090469e-02
#> Locus_1460 -4.197434e-02 -2.500077e-02 -3.352395e-02 -2.991172e-02
#> Locus_1461 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1463 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1468 -8.942676e-02 -4.409562e-02 -3.620539e-02 1.759536e-02
#> Locus_1470 -5.761153e-03 -5.611912e-02 -5.124809e-02 -6.394950e-02
#> Locus_1477 -3.425117e-02 -6.768357e-02 -1.326821e-02 1.169831e-01
#> Locus_1479 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1482 1.441861e-02 -1.389069e-02 -1.015649e-02 1.526350e-03
#> Locus_1485 -6.563856e-02 3.966337e-03 -1.287850e-02 -1.552423e-02
#> Locus_1486 2.972787e-02 -2.048425e-02 1.849201e-02 1.846905e-02
#> Locus_1487 -3.204999e-02 1.363418e-02 1.191967e-02 1.726288e-02
#> Locus_1488 -4.042243e-03 2.632964e-02 1.390425e-02 -1.343328e-02
#> Locus_1489 2.443710e-02 -1.821699e-03 -7.522013e-03 3.766558e-03
#> Locus_1490 5.647232e-02 -1.218385e-03 2.002784e-02 -2.791865e-02
#> Locus_1494 -1.623746e-02 -5.331885e-04 8.141886e-03 1.446149e-02
#> Locus_1496 -7.069997e-02 7.254876e-02 5.480329e-02 2.525171e-02
#> Locus_1500 -1.028249e-01 4.966632e-03 -4.323366e-02 -1.130891e-02
#> Locus_1502 1.248905e-02 6.865864e-02 3.191644e-03 5.196264e-03
#> Locus_1506 4.576198e-02 -2.991634e-02 -2.039307e-02 3.113941e-02
#> Locus_1508 5.096541e-02 3.722396e-02 -6.792043e-02 -1.275150e-02
#> Locus_1509 -1.118799e-02 1.955435e-02 7.455765e-02 -1.694437e-02
#> Locus_1514 2.717725e-02 -4.613789e-03 5.208706e-03 2.898540e-02
#> Locus_1515 3.383193e-02 -3.495956e-02 3.242889e-03 -7.642903e-04
#> Locus_1517 -8.577003e-03 1.541952e-02 2.044963e-02 2.157980e-02
#> Locus_1520 5.224601e-02 1.752089e-02 -5.821682e-03 2.630406e-02
#> Locus_1523 -2.296370e-02 -1.170472e-02 2.881655e-02 -1.239734e-02
#> Locus_1524 5.209729e-05 7.880453e-02 -7.065796e-02 5.258539e-02
#> Locus_1529 2.111377e-02 -1.939056e-02 -2.304468e-02 5.357261e-02
#> Locus_1534 -2.839484e-02 -9.535501e-03 -3.256073e-02 2.623626e-02
#> Locus_1536 -2.003401e-02 2.096491e-02 6.859537e-02 4.525872e-02
#> Locus_1540 -2.363913e-02 8.945338e-03 6.407780e-02 1.838616e-02
#> Locus_1545 4.776939e-02 1.319064e-01 1.064963e-02 1.764275e-01
#> Locus_1549 3.668647e-04 -1.058296e-02 -2.221997e-04 -3.031919e-03
#> Locus_1551 -6.970334e-02 3.516146e-03 -2.949983e-02 6.252167e-03
#> Locus_1554 9.930983e-02 -3.516357e-03 1.134482e-02 1.176042e-01
#> Locus_1555 -5.710411e-02 6.931152e-04 -2.770838e-02 -5.240405e-03
#> Locus_1561 3.329919e-02 -3.580551e-02 1.642082e-02 -1.230606e-01
#> Locus_1566 -1.200595e-02 -6.030078e-03 -1.997774e-02 2.836335e-02
#> Locus_1572 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1573 -6.950729e-03 -8.270729e-03 1.591347e-02 -1.775527e-02
#> Locus_1576 4.888559e-02 4.797514e-02 -4.108675e-02 1.707978e-02
#> Locus_1586 2.099629e-03 -1.627395e-02 1.494820e-02 -3.814368e-02
#> Locus_1587 3.319126e-02 -9.811668e-03 -1.730381e-02 -2.813140e-02
#> Locus_1590 -4.062883e-03 -2.320239e-02 -1.282688e-02 3.263786e-02
#> Locus_1591 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1596 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1597 4.914566e-02 -1.173032e-01 -9.726057e-02 -9.821434e-02
#> Locus_1598 4.935799e-02 1.348748e-03 -1.625878e-02 -1.028388e-02
#> Locus_1601 -2.658842e-02 -6.684867e-02 5.327379e-02 5.994099e-02
#> Locus_1602 2.334647e-03 7.577779e-04 1.560524e-02 -1.089229e-03
#> Locus_1603 3.163515e-02 -2.294814e-02 -1.997833e-02 -1.632825e-02
#> Locus_1604 -4.064502e-02 4.450591e-03 -2.602028e-02 1.294104e-01
#> Locus_1611 -3.640436e-02 9.804688e-03 2.449487e-02 -4.080166e-02
#> Locus_1612 -6.723920e-03 1.815012e-02 -8.368509e-03 3.840499e-02
#> Locus_1614 -7.318880e-03 -3.607567e-02 -1.959786e-02 4.442494e-02
#> Locus_1618 3.338327e-02 2.102971e-03 -8.776801e-03 -7.078292e-06
#> Locus_1619 4.063201e-02 -1.106635e-01 -4.834738e-02 -1.529894e-01
#> Locus_1620 5.678857e-02 4.508696e-02 3.332466e-02 1.526859e-01
#> Locus_1621 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1624 -2.469984e-02 5.342878e-04 8.181925e-03 4.181347e-04
#> Locus_1626 1.326399e-03 -1.875669e-03 1.576252e-02 -2.066468e-02
#> Locus_1632 7.933021e-03 1.005313e-02 -1.360238e-02 6.886742e-02
#> Locus_1633 -2.971258e-02 -4.713822e-02 5.896398e-02 4.740113e-02
#> Locus_1635 -3.823109e-02 -2.355878e-02 1.173724e-02 -1.235119e-02
#> Locus_1636 -8.358911e-02 -2.646945e-03 -3.935238e-02 2.227388e-02
#> Locus_1641 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1644 5.426820e-02 9.323286e-02 -2.881580e-02 -7.892617e-02
#> Locus_1645 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1646 4.877633e-03 -3.933227e-02 1.321871e-02 4.214035e-02
#> Locus_1647 -2.529188e-02 -6.515338e-04 -1.844710e-02 1.548470e-03
#> Locus_1648 -5.871846e-02 2.916115e-02 1.092667e-03 4.060146e-02
#> Locus_1649 3.415818e-03 -9.053577e-03 6.493709e-02 -8.939615e-02
#> Locus_1652 1.102251e-01 -5.955781e-02 -2.546857e-02 -4.721991e-02
#> Locus_1656 -6.085002e-03 -2.946494e-02 -9.410080e-03 -1.976686e-02
#> Locus_1658 -1.617028e-02 1.115959e-02 -3.293421e-02 1.845130e-02
#> Locus_1660 -1.316809e-01 6.364592e-03 -1.101057e-02 -3.102597e-02
#> Locus_1661 -1.424124e-02 -2.225207e-02 3.874270e-02 -9.301530e-03
#> Locus_1664 1.323214e-02 6.399571e-03 -2.071748e-02 2.026937e-02
#> Locus_1665 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1670 2.568139e-02 -1.128866e-02 -7.822825e-04 -1.142212e-02
#> Locus_1681 1.762852e-02 3.058523e-03 -1.294996e-02 1.970524e-02
#> Locus_1682 -6.370765e-02 3.156033e-03 1.747317e-02 -1.659098e-02
#> Locus_1687 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1690 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1691 3.919380e-02 9.388675e-03 1.695662e-02 5.407071e-02
#> Locus_1692 3.557513e-04 -1.653274e-02 -6.033303e-03 8.077438e-03
#> Locus_1693 1.504163e-02 9.515239e-02 2.779748e-02 1.053633e-01
#> Locus_1694 1.955232e-02 -3.718909e-02 -1.942255e-02 6.017189e-02
#> Locus_1697 7.443418e-02 -4.347381e-02 6.944188e-03 3.868109e-02
#> Locus_1699 7.737697e-02 -6.149311e-02 -1.368563e-02 -3.303808e-03
#> Locus_1702 3.563963e-02 -5.681266e-03 5.168667e-03 4.302876e-02
#> Locus_1703 1.075676e-02 -1.312161e-02 -1.216288e-02 -3.130129e-02
#> Locus_1708 4.196890e-02 -2.733331e-02 1.085746e-02 1.491760e-02
#> Locus_1713 -7.384783e-03 1.428675e-02 2.788152e-02 3.346286e-02
#> Locus_1715 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1724 1.022711e-02 7.306866e-03 4.016829e-02 -1.094338e-01
#> Locus_1728 3.135668e-02 1.659961e-02 5.934113e-02 1.249024e-01
#> Locus_1730 2.943372e-02 1.625551e-02 1.164862e-02 3.883238e-02
#> Locus_1734 1.055373e-02 1.542696e-02 8.361247e-03 -1.554663e-02
#> Locus_1737 4.151976e-05 -4.813638e-02 1.206148e-03 9.225721e-02
#> Locus_1742 1.574760e-04 -5.266560e-02 3.236223e-02 3.120712e-02
#> Locus_1745 -6.157383e-02 4.205860e-02 -2.898008e-02 1.791084e-02
#> Locus_1748 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1750 2.476349e-02 9.455238e-03 5.482704e-04 1.057840e-01
#> Locus_1751 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1755 3.443307e-02 -1.811489e-02 -1.964367e-02 -3.570921e-02
#> Locus_1756 -1.546256e-03 -1.246929e-02 -5.940525e-03 1.616547e-02
#> Locus_1757 1.781294e-02 3.563578e-02 1.382199e-02 6.201326e-02
#> Locus_1774 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1776 2.110842e-02 -6.324631e-02 3.283648e-02 5.187576e-02
#> Locus_1777 2.741914e-02 -2.117716e-03 4.601331e-03 2.007691e-02
#> Locus_1778 -6.883828e-02 3.362353e-03 -2.899179e-02 -4.091415e-03
#> Locus_1783 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1784 7.774037e-02 1.667205e-02 -2.106484e-03 1.818804e-01
#> Locus_1786 2.750045e-02 2.217667e-03 5.694019e-03 1.804796e-02
#> Locus_1787 -8.210139e-03 4.836564e-03 2.022743e-02 1.854788e-02
#> Locus_1797 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1799 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1801 3.500462e-02 5.290265e-03 -1.395426e-02 -7.205694e-02
#> Locus_1804 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1805 -3.883290e-02 -5.686298e-02 -3.356403e-02 -4.163281e-02
#> Locus_1807 7.156945e-02 -1.580443e-02 1.936962e-03 1.350245e-02
#> Locus_1808 -2.621198e-03 -7.475565e-02 1.359488e-02 -9.379353e-03
#> Locus_1811 3.099369e-02 6.192089e-03 4.379684e-03 4.671845e-02
#> Locus_1813 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1816 -8.577003e-03 1.541952e-02 2.044963e-02 2.157980e-02
#> Locus_1822 -7.849148e-02 -5.256365e-04 -4.015879e-02 -2.395047e-02
#> Locus_1823 4.351219e-02 -2.033756e-02 -4.140468e-02 1.087926e-01
#> Locus_1826 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1828 -1.835452e-02 2.560846e-02 1.432856e-02 1.257867e-02
#> Locus_1833 2.300792e-02 -4.295687e-02 4.805865e-03 6.587919e-02
#> Locus_1839 -2.210673e-02 -2.348236e-02 -3.265738e-02 -1.675076e-02
#> Locus_1840 -1.413199e-02 1.352250e-02 -1.253111e-02 4.987254e-02
#> Locus_1841 -2.036577e-02 -9.234659e-02 2.779177e-02 5.202809e-02
#> Locus_1842 2.813787e-02 -7.667658e-02 2.875812e-02 -9.571185e-02
#> Locus_1843 3.599593e-02 -1.070404e-02 4.310342e-03 9.337575e-03
#> Locus_1844 2.990097e-02 -9.909722e-02 3.656622e-02 1.750249e-01
#> Locus_1846 1.097996e-02 6.745721e-03 -9.956802e-03 -1.151081e-02
#> Locus_1848 -2.885598e-02 1.397960e-03 3.222308e-02 -1.971706e-02
#> Locus_1849 -4.334569e-02 4.509186e-02 -1.481324e-02 2.182959e-02
#> Locus_1852 -1.120429e-01 1.135109e-02 2.246550e-02 -3.694643e-02
#> Locus_1853 -8.667821e-03 -3.637613e-02 2.558670e-02 -7.916256e-02
#> Locus_1856 2.568139e-02 -1.128866e-02 -7.822825e-04 -1.142212e-02
#> Locus_1861 3.444771e-02 -6.796609e-03 -1.625855e-03 1.357374e-02
#> Locus_1863 -3.483420e-03 -1.177307e-02 3.720931e-03 -1.372479e-02
#> Locus_1868 3.230829e-02 7.974142e-02 -8.902200e-03 -5.590455e-02
#> Locus_1870 2.334647e-03 7.577779e-04 1.560524e-02 -1.089229e-03
#> Locus_1871 3.021946e-02 2.867185e-02 1.478528e-02 3.999855e-03
#> Locus_1872 3.185745e-02 -6.264586e-03 -5.778674e-03 2.524380e-02
#> Locus_1873 1.172720e-01 -3.704290e-02 1.294700e-02 2.169471e-02
#> Locus_1881 -4.665565e-02 1.955466e-02 -5.489244e-02 -7.094527e-03
#> Locus_1882 -3.467309e-03 3.502344e-03 1.219254e-02 -4.030481e-03
#> Locus_1887 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1894 -1.626090e-02 -1.393242e-02 -5.731148e-02 1.017099e-01
#> Locus_1896 3.163515e-02 -2.294814e-02 -1.997833e-02 -1.632825e-02
#> Locus_1908 -9.558054e-02 5.273754e-02 8.840474e-04 -5.483241e-03
#> Locus_1911 -5.311546e-03 -5.102429e-02 2.075980e-02 2.310608e-02
#> Locus_1913 9.614168e-03 9.997668e-03 3.944643e-02 -1.338734e-03
#> Locus_1919 -6.723920e-03 1.815012e-02 -8.368509e-03 3.840499e-02
#> Locus_1921 -1.544864e-03 -1.627300e-02 2.240126e-02 6.845729e-03
#> Locus_1932 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1933 -1.328504e-02 1.973678e-02 -9.785433e-03 3.016933e-02
#> Locus_1935 -2.666289e-02 2.066910e-03 -1.474387e-02 -2.099082e-03
#> Locus_1939 5.111026e-03 1.055836e-02 1.891968e-02 1.028314e-02
#> Locus_1941 2.114486e-04 2.609162e-02 3.194321e-02 1.075909e-02
#> Locus_1942 1.597473e-02 -7.542226e-04 -7.481974e-03 -1.027680e-02
#> Locus_1945 4.164830e-02 -6.120153e-03 4.127459e-02 -1.436422e-02
#> Locus_1948 2.921811e-02 -1.764555e-02 -1.022962e-02 3.886995e-02
#> Locus_1949 7.190535e-02 -6.535793e-02 -3.833952e-02 1.858091e-02
#> Locus_1950 -3.075734e-02 3.586240e-02 3.504893e-02 1.008828e-02
#> Locus_1951 3.422588e-02 -4.764844e-03 8.200465e-03 -2.854771e-03
#> Locus_1952 4.216538e-02 2.054419e-02 9.240534e-03 4.331352e-02
#> Locus_1953 -2.119859e-02 2.452926e-02 6.866149e-02 -2.675156e-02
#> Locus_1958 2.235116e-02 -1.367006e-02 -1.221045e-02 4.930070e-03
#> Locus_1961 1.139101e-01 4.251395e-02 1.566094e-02 -2.370489e-02
#> Locus_1962 5.167670e-02 -3.468472e-02 3.363739e-02 -8.677481e-02
#> Locus_1965 2.033211e-02 2.500959e-02 -3.705946e-02 -2.413451e-03
#> Locus_1967 -5.332579e-02 4.133821e-03 -2.948774e-02 -4.198165e-03
#> Locus_1968 2.114486e-04 2.609162e-02 3.194321e-02 1.075909e-02
#> Locus_1971 -1.788643e-02 -2.193911e-02 4.909915e-02 -7.821047e-03
#> Locus_1974 1.625802e-02 -1.029425e-03 -3.319832e-02 -1.430058e-02
#> Locus_1977 2.649132e-02 1.054069e-02 -4.883764e-03 -6.454852e-02
#> Locus_1982 -3.023986e-04 -4.741815e-03 -4.607061e-02 -2.995503e-02
#> Locus_1983 -1.134938e-01 -5.134392e-02 -2.765277e-02 1.098722e-01
#> Locus_1984 -8.979646e-02 8.158219e-03 -1.998991e-02 5.044954e-03
#> Locus_1988 -4.698956e-03 4.178968e-02 1.965541e-03 1.651698e-03
#> Locus_1990 4.732985e-02 1.948826e-02 5.911929e-02 9.681807e-02
#> Locus_1991 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_1993 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2003 4.215029e-02 1.450672e-02 -2.341254e-02 4.593810e-02
#> Locus_2007 -5.332579e-02 4.133821e-03 -2.948774e-02 -4.198165e-03
#> Locus_2014 -1.881951e-02 -7.987730e-02 3.373229e-02 3.586261e-02
#> Locus_2016 5.541807e-02 4.757827e-02 -3.223923e-02 -6.012056e-02
#> Locus_2017 1.373040e-02 1.018152e-04 3.307657e-02 5.879629e-02
#> Locus_2019 1.393756e-02 2.975029e-02 5.652124e-02 1.023250e-02
#> Locus_2020 -2.303316e-02 -1.449832e-02 -1.094004e-02 -7.252869e-03
#> Locus_2021 -2.065657e-02 3.178165e-02 -4.973374e-02 -1.371906e-02
#> Locus_2027 1.781294e-02 3.563578e-02 1.382199e-02 6.201326e-02
#> Locus_2030 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2033 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2036 -3.352527e-02 -1.175957e-04 1.012600e-03 -1.753860e-02
#> Locus_2041 -6.563856e-02 3.966337e-03 -1.287850e-02 -1.552423e-02
#> Locus_2045 3.738079e-02 -4.208833e-02 2.857132e-03 1.911104e-02
#> Locus_2048 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2050 3.422287e-02 -4.721163e-02 3.816820e-02 -7.594499e-02
#> Locus_2053 1.653642e-02 4.577760e-02 1.165123e-01 -1.599171e-01
#> Locus_2055 -1.401632e-02 -1.006366e-02 -1.944670e-02 8.266316e-02
#> Locus_2060 -1.000906e-02 5.792617e-03 -5.277030e-02 -2.880971e-02
#> Locus_2063 -5.749339e-04 2.282730e-02 1.711713e-03 -9.402803e-03
#> Locus_2064 -5.738787e-02 -1.182708e-01 4.872925e-02 -3.182623e-02
#> Locus_2068 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2069 -8.551433e-03 -4.862215e-02 2.246042e-02 4.458747e-02
#> Locus_2080 3.624394e-02 2.652587e-02 -2.853245e-02 -1.779130e-02
#> Locus_2085 -3.741650e-03 -4.261579e-03 -8.199546e-04 -8.718234e-03
#> Locus_2086 -1.217000e-02 -5.892989e-02 -1.882016e-02 -3.953373e-02
#> Locus_2087 -6.085002e-03 -2.946494e-02 -9.410080e-03 -1.976686e-02
#> Locus_2091 8.370259e-03 -8.500667e-03 -1.693862e-02 -4.053514e-02
#> Locus_2094 6.767420e-02 7.993692e-04 1.899413e-03 -6.966072e-03
#> Locus_2102 4.355797e-02 4.928198e-02 -5.077055e-03 -1.931105e-02
#> Locus_2106 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2108 -5.311546e-03 -5.102429e-02 2.075980e-02 2.310608e-02
#> Locus_2109 8.276465e-04 -1.373907e-02 6.658967e-03 2.527783e-02
#> Locus_2113 5.046014e-02 7.044883e-03 -5.083586e-02 1.451193e-02
#> Locus_2115 2.567891e-02 -1.012492e-02 -9.861874e-03 -3.811248e-03
#> Locus_2120 -9.767508e-03 -1.358251e-02 8.475807e-03 4.582271e-02
#> Locus_2128 -2.577926e-02 -1.277031e-02 6.502598e-03 -4.119007e-02
#> Locus_2131 -2.088981e-02 -2.865671e-02 1.327638e-02 -4.200780e-02
#> Locus_2133 -5.859968e-02 6.875080e-02 -2.581048e-02 -1.194927e-02
#> Locus_2135 1.441861e-02 -1.389069e-02 -1.015649e-02 1.526350e-03
#> Locus_2136 -3.429253e-02 2.449427e-02 -3.615254e-03 -3.549423e-02
#> Locus_2137 -1.390146e-02 -1.654146e-02 3.182694e-02 -3.551054e-02
#> Locus_2138 3.919380e-02 9.388675e-03 1.695662e-02 5.407071e-02
#> Locus_2139 -4.124022e-03 1.896239e-02 2.538280e-04 1.105450e-02
#> Locus_2140 6.304317e-05 1.270028e-02 1.155616e-02 -3.107421e-03
#> Locus_2145 -6.342591e-02 1.120635e-03 -2.527999e-02 3.057247e-03
#> Locus_2148 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2154 1.108902e-02 5.378590e-02 5.453479e-03 1.004182e-01
#> Locus_2157 2.463392e-02 1.295164e-02 6.403378e-03 -3.475730e-02
#> Locus_2159 -1.385421e-02 -3.988770e-02 8.000611e-03 4.898559e-04
#> Locus_2162 -1.508647e-02 3.279726e-02 -2.993597e-02 -1.082637e-01
#> Locus_2165 4.605376e-02 -3.683884e-02 -3.013482e-02 -1.480190e-02
#> Locus_2166 -2.225244e-02 -1.064958e-03 -1.295846e-02 -8.366488e-03
#> Locus_2168 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2179 -7.122920e-02 -9.664717e-03 -3.525472e-02 -2.647913e-03
#> Locus_2180 -4.249920e-03 -4.891462e-02 -2.393026e-03 2.597357e-02
#> Locus_2181 -7.865501e-03 -1.764595e-02 8.383029e-03 3.773468e-02
#> Locus_2183 -1.671963e-02 -1.972570e-03 1.821873e-02 -1.676802e-01
#> Locus_2185 -3.604924e-02 -4.882284e-03 -6.062544e-02 -6.255459e-02
#> Locus_2186 -1.116153e-03 -2.054736e-03 -6.270666e-04 1.313752e-02
#> Locus_2188 -2.095819e-02 4.795867e-03 9.001880e-03 9.136369e-03
#> Locus_2191 -1.225392e-02 6.224463e-03 8.354467e-03 1.427123e-02
#> Locus_2203 -4.584738e-03 -5.306474e-03 2.203977e-03 1.728969e-02
#> Locus_2208 -7.174667e-02 1.223347e-02 1.964158e-02 -2.293463e-02
#> Locus_2211 -1.738170e-01 6.794839e-02 -1.938883e-02 1.903459e-02
#> Locus_2213 -8.658831e-03 -3.134549e-02 6.065116e-03 5.402565e-03
#> Locus_2215 -4.229935e-02 -4.893694e-02 -1.981372e-02 -6.193752e-02
#> Locus_2216 -2.693429e-02 -1.717971e-02 1.295580e-02 5.337279e-03
#> Locus_2219 -4.850605e-02 -5.522424e-02 -7.109875e-02 2.089521e-02
#> Locus_2223 -1.094758e-02 -1.790432e-04 1.489804e-02 1.894356e-02
#> Locus_2228 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2229 1.465694e-02 -8.096988e-03 -5.233872e-03 3.046243e-02
#> Locus_2230 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2231 8.462378e-03 -1.067476e-03 -4.003902e-05 1.404336e-02
#> Locus_2234 1.001060e-02 -4.974910e-03 5.896158e-03 9.807186e-03
#> Locus_2236 3.745051e-02 -5.101485e-03 -1.343113e-02 6.982863e-02
#> Locus_2238 1.099126e-02 3.287421e-03 -5.962069e-03 -1.141723e-02
#> Locus_2243 -7.607472e-02 1.113658e-02 -1.846331e-02 -7.580468e-03
#> Locus_2246 -2.586562e-02 -4.437765e-02 2.703283e-02 -9.095738e-02
#> Locus_2250 -7.965356e-02 -7.770102e-03 -1.524890e-02 -3.139345e-02
#> Locus_2251 8.468130e-03 3.880943e-03 5.109097e-02 -2.242328e-02
#> Locus_2254 3.764093e-03 -1.187866e-01 -5.935774e-03 -7.462280e-02
#> Locus_2260 -6.507428e-03 -1.683132e-02 -7.352437e-03 -2.750482e-02
#> Locus_2261 5.019760e-02 -4.796585e-02 3.068622e-02 -8.622178e-02
#> Locus_2267 4.840417e-02 -2.524117e-03 1.553083e-02 2.795989e-01
#> Locus_2268 -5.167654e-02 -9.570841e-03 -6.172658e-02 1.828187e-02
#> Locus_2271 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2273 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2274 7.338478e-02 3.926992e-02 -7.375509e-02 -2.198714e-03
#> Locus_2275 2.009057e-02 -1.024798e-02 -4.118544e-02 5.261646e-03
#> Locus_2277 1.596389e-02 6.518177e-03 3.024597e-02 8.790839e-02
#> Locus_2282 -5.113699e-02 -5.919508e-02 9.181417e-02 2.246693e-01
#> Locus_2287 1.873616e-02 8.063262e-02 3.365538e-02 -2.913365e-03
#> Locus_2294 -3.883290e-02 -5.686298e-02 -3.356403e-02 -4.163281e-02
#> Locus_2298 -1.105337e-02 -1.174118e-02 -1.632869e-02 -8.375382e-03
#> Locus_2305 -4.141735e-03 -2.704718e-02 -2.494750e-02 1.949926e-02
#> Locus_2308 -6.277569e-03 2.539342e-02 -3.036149e-02 -3.727778e-02
#> Locus_2309 -4.062883e-03 -2.320239e-02 -1.282688e-02 3.263786e-02
#> Locus_2312 8.479475e-03 8.538094e-03 -9.132195e-03 4.577443e-02
#> Locus_2318 5.172520e-02 2.358191e-02 -6.850255e-02 -1.325868e-02
#> Locus_2319 1.649764e-02 5.035510e-02 6.088487e-02 3.836769e-02
#> Locus_2321 -4.971139e-03 -6.728694e-02 -1.152600e-01 -3.237959e-02
#> Locus_2322 -3.768343e-02 -8.207028e-02 -5.081206e-02 1.799460e-01
#> Locus_2323 3.451998e-02 -2.074862e-02 1.778927e-02 -2.770277e-02
#> Locus_2325 2.052725e-02 -3.265939e-02 8.402118e-02 -1.273686e-01
#> Locus_2326 -6.319245e-03 -5.176659e-03 1.432355e-02 2.156920e-02
#> Locus_2327 -2.595072e-02 -3.580169e-02 3.160695e-02 -1.058550e-01
#> Locus_2333 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2336 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2338 -1.056629e-03 1.827291e-01 -7.822796e-02 -8.042416e-02
#> Locus_2339 -1.094758e-02 -1.790432e-04 1.489804e-02 1.894356e-02
#> Locus_2344 -4.249168e-02 -5.907196e-02 -8.362635e-03 5.801484e-02
#> Locus_2345 -3.467309e-03 3.502344e-03 1.219254e-02 -4.030481e-03
#> Locus_2348 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2349 2.883722e-02 -2.778139e-02 -2.031299e-02 3.052700e-03
#> Locus_2354 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2359 1.234525e-02 -4.217132e-03 2.150140e-02 8.717957e-03
#> Locus_2361 6.128118e-02 -2.238048e-02 -1.152505e-02 2.970461e-02
#> Locus_2364 8.084210e-02 3.268522e-03 5.823122e-02 3.970649e-02
#> Locus_2365 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2366 -5.991047e-02 2.346073e-02 -2.649787e-02 1.787793e-02
#> Locus_2368 -7.417503e-02 8.672815e-03 8.377961e-03 5.583091e-02
#> Locus_2371 -6.970334e-02 3.516146e-03 -2.949983e-02 6.252167e-03
#> Locus_2375 5.526346e-02 3.053430e-02 3.908593e-04 1.318419e-01
#> Locus_2381 2.921811e-02 -1.764555e-02 -1.022962e-02 3.886995e-02
#> Locus_2388 -3.604250e-02 -3.197392e-03 -2.595696e-02 -2.689620e-03
#> Locus_2394 1.780534e-02 4.395921e-02 -8.904834e-02 -2.310206e-02
#> Locus_2400 5.667839e-04 -5.619264e-02 -3.229260e-03 2.600923e-02
#> Locus_2402 -5.411583e-02 -2.003845e-02 -3.763556e-02 1.245940e-01
#> Locus_2406 2.921834e-02 -4.431532e-02 1.178304e-02 -9.034984e-02
#> Locus_2413 7.496536e-03 -5.027352e-03 -6.235871e-03 -2.523560e-02
#> Locus_2414 -3.352527e-02 -1.175957e-04 1.012600e-03 -1.753860e-02
#> Locus_2415 -3.232866e-02 -9.675715e-03 -2.820765e-02 -2.608285e-02
#> Locus_2421 -2.723783e-02 2.489421e-02 -1.303215e-02 -1.150189e-02
#> Locus_2424 -6.736870e-02 4.273924e-03 -1.389459e-02 5.162938e-03
#> Locus_2425 -1.545344e-02 -1.124576e-01 -8.784160e-04 2.897757e-02
#> Locus_2426 -5.744981e-02 2.309621e-02 -2.923391e-02 6.856337e-03
#> Locus_2431 4.715822e-02 -6.584488e-03 -2.871598e-02 6.567608e-02
#> Locus_2432 -6.883828e-02 3.362353e-03 -2.899179e-02 -4.091415e-03
#> Locus_2433 -2.715724e-03 -4.914625e-02 3.538715e-03 5.810178e-02
#> Locus_2434 -5.332579e-02 4.133821e-03 -2.948774e-02 -4.198165e-03
#> Locus_2435 -3.483420e-03 -1.177307e-02 3.720931e-03 -1.372479e-02
#> Locus_2436 3.334534e-02 1.394351e-02 -9.371678e-02 3.959804e-02
#> Locus_2437 -5.311546e-03 -5.102429e-02 2.075980e-02 2.310608e-02
#> Locus_2439 -3.915015e-04 1.753582e-02 1.600614e-03 -1.091876e-02
#> Locus_2448 5.261601e-02 -5.963318e-02 -4.872132e-02 5.854554e-02
#> Locus_2449 -4.806669e-02 -9.299567e-03 -1.386230e-02 -8.228734e-02
#> Locus_2450 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2451 3.461969e-02 -4.833636e-02 -2.079821e-02 6.116284e-03
#> Locus_2452 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2457 -3.149331e-02 2.778068e-02 -6.747568e-02 -1.239218e-02
#> Locus_2461 2.075988e-05 -2.406819e-02 6.030742e-04 4.612860e-02
#> Locus_2463 2.737506e-02 -3.993551e-02 6.109469e-02 -1.016568e-01
#> Locus_2466 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2467 -4.080430e-03 -3.265438e-02 -1.122548e-02 4.570395e-02
#> Locus_2473 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2475 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2477 1.093517e-01 -1.294787e-01 -2.534206e-02 2.622623e-01
#> Locus_2480 -7.846414e-03 -4.288471e-02 1.285927e-02 8.380496e-02
#> Locus_2494 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2497 2.527303e-03 -1.349807e-02 4.256249e-03 -7.629282e-03
#> Locus_2501 7.102376e-03 -1.781320e-02 -3.368620e-02 -6.678220e-02
#> Locus_2502 3.194945e-02 -1.508445e-03 -1.496395e-02 -2.055359e-02
#> Locus_2507 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2510 -1.299424e-02 6.179418e-03 3.468041e-02 3.506097e-02
#> Locus_2512 -2.359245e-02 -2.014947e-02 -4.820700e-02 7.255526e-02
#> Locus_2514 -2.015244e-02 -1.722151e-02 -3.049837e-02 -3.543272e-02
#> Locus_2517 -1.185264e-02 -1.647150e-02 2.792727e-02 4.818788e-02
#> Locus_2518 -2.638929e-02 6.905968e-04 -1.478996e-02 1.716944e-02
#> Locus_2519 5.669294e-02 -4.978126e-02 -5.084699e-02 9.349783e-02
#> Locus_2522 -8.754187e-03 -6.798346e-02 4.611693e-02 -1.289299e-01
#> Locus_2526 -6.925319e-03 -7.168252e-02 8.648011e-03 6.480763e-02
#> Locus_2529 7.837253e-03 8.601560e-03 -1.336425e-02 4.681251e-02
#> Locus_2530 -9.815558e-02 6.482188e-03 -1.202317e-02 -1.348737e-02
#> Locus_2532 2.158563e-02 -1.270599e-02 -2.016940e-02 1.998192e-02
#> Locus_2540 4.899300e-03 1.860631e-02 -1.680826e-02 1.623949e-02
#> Locus_2542 3.661046e-03 -1.117891e-03 3.136776e-02 -2.175391e-02
#> Locus_2546 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2550 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2555 -5.279326e-03 -2.047345e-02 3.770301e-02 4.249470e-02
#> Locus_2559 -2.720030e-02 1.025090e-03 9.981146e-02 -4.841892e-02
#> Locus_2561 6.254830e-02 5.714619e-02 -1.742520e-02 1.167825e-01
#> Locus_2565 1.711144e-02 -2.360582e-02 1.908410e-02 -3.797249e-02
#> Locus_2566 -5.332579e-02 4.133821e-03 -2.948774e-02 -4.198165e-03
#> Locus_2567 5.569228e-02 5.503298e-02 -4.947786e-02 -4.760043e-02
#> Locus_2572 -8.125766e-03 -4.640478e-02 -2.565375e-02 6.527572e-02
#> Locus_2573 1.107060e-01 -8.787319e-03 -7.164107e-03 -9.176059e-03
#> Locus_2577 -3.463456e-03 -2.280328e-02 4.129621e-03 1.031527e-02
#> Locus_2579 1.209515e-01 6.261191e-02 7.880581e-02 1.204457e-01
#> Locus_2581 -3.231515e-02 -3.650418e-02 -1.701364e-02 -9.158135e-02
#> Locus_2583 4.810288e-02 -3.628821e-02 2.810669e-02 1.938159e-01
#> Locus_2588 -2.366970e-02 -7.083806e-02 5.095904e-02 6.503855e-02
#> Locus_2589 -3.611032e-02 -3.243534e-02 8.380223e-03 -1.961651e-01
#> Locus_2592 6.011264e-02 3.715770e-02 5.933882e-02 5.126218e-02
#> Locus_2597 -2.225244e-02 -1.064958e-03 -1.295846e-02 -8.366488e-03
#> Locus_2598 1.474894e-02 -2.507142e-05 -3.346547e-02 -3.687801e-03
#> Locus_2599 -6.085002e-03 -2.946494e-02 -9.410080e-03 -1.976686e-02
#> Locus_2601 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2602 1.955670e-02 1.754538e-02 1.950131e-02 -7.260948e-02
#> Locus_2611 -1.858533e-02 1.139254e-02 6.550209e-02 9.469807e-03
#> Locus_2616 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2617 -3.078872e-02 -8.674717e-06 -7.586947e-03 -2.712449e-02
#> Locus_2619 2.741914e-02 -2.117716e-03 4.601331e-03 2.007691e-02
#> Locus_2620 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2629 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2632 5.912333e-03 6.467415e-02 4.961284e-02 3.848222e-03
#> Locus_2634 -2.970913e-02 -3.098829e-03 -5.266334e-02 -1.899684e-02
#> Locus_2636 1.112217e-02 3.015833e-03 -8.486391e-03 -4.882448e-02
#> Locus_2639 2.848947e-02 5.885215e-02 4.453450e-02 2.855331e-02
#> Locus_2644 2.948243e-02 -3.989576e-02 -2.704767e-02 3.813893e-03
#> Locus_2647 1.555828e-02 1.163915e-02 2.225216e-02 -2.376152e-02
#> Locus_2648 -4.983461e-03 4.041644e-03 1.519906e-03 -1.140428e-03
#> Locus_2652 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2655 1.729487e-02 -2.889729e-02 1.897300e-02 -3.948845e-02
#> Locus_2656 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2657 4.280187e-02 5.260175e-02 -2.271455e-02 -1.056222e-02
#> Locus_2660 4.780188e-02 -1.178772e-02 -1.893330e-02 1.519272e-03
#> Locus_2662 -4.679571e-02 8.191213e-02 -2.837088e-02 -7.484423e-02
#> Locus_2665 6.822320e-02 1.238363e-01 -6.120072e-02 1.807003e-01
#> Locus_2668 -7.322863e-02 -6.953724e-02 -5.495073e-02 5.095432e-02
#> Locus_2669 -5.261471e-02 3.859663e-02 4.227968e-03 -1.985531e-02
#> Locus_2670 -6.487255e-02 5.319962e-02 -4.595700e-03 -2.825829e-02
#> Locus_2671 -6.085002e-03 -2.946494e-02 -9.410080e-03 -1.976686e-02
#> Locus_2675 6.786082e-02 -5.917233e-02 -2.645166e-02 -5.562958e-02
#> Locus_2677 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2685 -3.127292e-02 -1.020077e-01 -1.013970e-02 -1.196769e-01
#> Locus_2687 1.971726e-02 -1.550934e-02 1.259585e-02 8.661866e-03
#> Locus_2696 7.223898e-02 -1.360942e-02 -2.645531e-02 5.285830e-03
#> Locus_2698 -2.300373e-02 -1.137283e-02 5.065775e-02 6.418911e-02
#> Locus_2699 -5.744981e-02 2.309621e-02 -2.923391e-02 6.856337e-03
#> Locus_2702 -1.483047e-02 -8.191746e-03 -2.957601e-03 2.274046e-02
#> Locus_2703 -2.788521e-03 -6.294507e-02 -1.413535e-02 -1.680164e-01
#> Locus_2705 -1.470913e-02 8.511136e-02 4.317515e-02 -1.792004e-02
#> Locus_2712 -1.373828e-02 -1.452289e-02 -8.622239e-03 5.830214e-03
#> Locus_2718 -5.035903e-03 -1.012240e-02 -2.278029e-02 -2.622109e-02
#> Locus_2719 -6.444441e-03 -7.397318e-04 3.542479e-02 7.321682e-03
#> Locus_2724 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2725 3.051396e-03 4.423140e-02 6.476099e-04 -9.749755e-03
#> Locus_2727 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2729 4.021137e-02 3.468878e-02 -4.538013e-03 1.172333e-01
#> Locus_2731 -3.274790e-02 -2.739803e-02 -2.415395e-02 -2.186595e-02
#> Locus_2733 6.033415e-02 7.279113e-02 -1.508230e-02 -6.740382e-02
#> Locus_2736 1.538882e-02 -2.328763e-02 1.663222e-03 3.368535e-02
#> Locus_2738 3.345816e-02 3.408683e-02 -1.232563e-02 -3.709894e-02
#> Locus_2739 4.885169e-02 -3.200559e-02 -2.980016e-02 -3.418286e-02
#> Locus_2753 3.433330e-02 7.222409e-04 -1.374905e-03 3.835643e-02
#> Locus_2755 2.436360e-02 -1.863142e-02 1.465820e-03 2.931711e-02
#> Locus_2758 -2.711241e-02 -6.272595e-02 5.700970e-02 6.484043e-02
#> Locus_2760 -8.919686e-02 -3.041226e-02 -2.042863e-02 -6.300619e-02
#> Locus_2761 -6.883828e-02 3.362353e-03 -2.899179e-02 -4.091415e-03
#> Locus_2764 3.345816e-02 3.408683e-02 -1.232563e-02 -3.709894e-02
#> Locus_2766 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2767 7.413657e-02 -2.071516e-02 -1.244847e-01 -1.757642e-03
#> Locus_2768 -1.755688e-03 3.923257e-02 -1.907561e-02 -9.080388e-03
#> Locus_2770 -3.491094e-02 3.999168e-02 -1.423621e-02 2.000992e-02
#> Locus_2774 -5.332579e-02 4.133821e-03 -2.948774e-02 -4.198165e-03
#> Locus_2776 -6.881032e-02 -4.441228e-02 -1.540877e-02 -9.095144e-03
#> Locus_2789 5.400175e-02 8.853378e-03 5.290365e-02 5.435121e-02
#> Locus_2792 2.435244e-02 -1.509160e-02 -4.581453e-03 -1.869975e-02
#> Locus_2793 -1.430234e-02 -1.503463e-02 2.299105e-02 -6.095375e-03
#> Locus_2794 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2795 -3.033977e-02 -4.482525e-02 -6.178551e-03 1.873334e-01
#> Locus_2796 1.374923e-02 -5.555101e-03 2.370703e-03 -2.188508e-02
#> Locus_2798 2.848947e-02 5.885215e-02 4.453450e-02 2.855331e-02
#> Locus_2804 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2806 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2808 1.485688e-02 -2.648643e-02 7.040219e-03 -1.738402e-02
#> Locus_2809 -4.124022e-03 1.896239e-02 2.538280e-04 1.105450e-02
#> Locus_2810 4.377929e-02 -2.719411e-02 -1.522536e-02 4.727748e-02
#> Locus_2811 4.870138e-03 6.789351e-02 1.298863e-02 -3.124893e-02
#> Locus_2814 2.855858e-02 4.962010e-02 6.978971e-04 -3.662299e-02
#> Locus_2816 -2.666289e-02 2.066910e-03 -1.474387e-02 -2.099082e-03
#> Locus_2817 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2818 -8.435842e-04 -7.002512e-02 -2.738970e-03 3.516944e-02
#> Locus_2823 4.339860e-02 -1.034087e-01 5.680177e-02 1.422416e-01
#> Locus_2830 -1.402277e-02 5.912754e-02 -1.502530e-02 6.740717e-02
#> Locus_2831 -1.155304e-01 1.076522e-01 6.275431e-02 -2.271399e-02
#> Locus_2832 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2835 1.283743e-02 -2.582387e-02 -1.961420e-02 9.567465e-02
#> Locus_2836 1.755673e-02 3.304345e-03 4.961542e-04 6.728358e-03
#> Locus_2839 1.182630e-01 -7.103831e-02 -1.879453e-01 -4.795258e-02
#> Locus_2842 -8.102434e-02 -4.030733e-02 -4.652353e-02 -4.524516e-02
#> Locus_2844 7.417382e-03 2.200909e-02 1.622521e-02 -1.690270e-02
#> Locus_2845 5.674550e-02 -1.372434e-02 5.168441e-02 -7.991006e-02
#> Locus_2850 -5.273890e-03 6.461698e-02 3.677255e-03 -7.751105e-03
#> Locus_2852 -8.084438e-03 -2.536783e-02 2.124665e-02 1.750845e-02
#> Locus_2854 -3.483420e-03 -1.177307e-02 3.720931e-03 -1.372479e-02
#> Locus_2856 -1.345440e-01 7.370275e-02 2.734523e-03 8.717736e-02
#> Locus_2861 -1.137844e-02 -4.547544e-02 5.854245e-02 -1.765544e-01
#> Locus_2865 1.097996e-02 6.745721e-03 -9.956802e-03 -1.151081e-02
#> Locus_2868 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2871 4.885169e-02 -3.200559e-02 -2.980016e-02 -3.418286e-02
#> Locus_2872 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2873 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2874 -8.952304e-02 1.740943e-02 -2.598295e-02 -1.515338e-02
#> Locus_2875 2.443710e-02 -1.821699e-03 -7.522013e-03 3.766558e-03
#> Locus_2878 3.708587e-02 -2.186623e-02 1.188137e-02 -7.703858e-02
#> Locus_2881 1.734956e-02 -1.628393e-02 6.807234e-03 -2.730972e-02
#> Locus_2884 -2.493286e-03 3.912642e-02 -8.424739e-03 -5.336894e-02
#> Locus_2890 2.567891e-02 -1.012492e-02 -9.861874e-03 -3.811248e-03
#> Locus_2895 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2896 -2.355635e-02 -7.256038e-02 2.574128e-02 -2.047681e-03
#> Locus_2904 -1.385994e-02 -6.467783e-02 3.303309e-02 5.674667e-02
#> Locus_2905 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2910 -1.059930e-02 1.226463e-02 3.433074e-02 -1.337578e-02
#> Locus_2915 -1.971856e-02 4.085491e-03 -2.400549e-02 4.873147e-02
#> Locus_2916 5.365858e-02 4.068251e-02 -6.135083e-02 1.167605e-02
#> Locus_2923 8.112120e-03 -3.877172e-02 -4.492877e-03 7.366583e-02
#> Locus_2932 -8.622061e-02 6.648342e-02 2.489779e-02 9.041173e-04
#> Locus_2933 1.871487e-02 -3.546313e-03 5.248745e-03 1.494205e-02
#> Locus_2934 -6.118433e-03 -6.135341e-02 1.470390e-02 4.395686e-02
#> Locus_2936 5.760986e-02 1.592862e-02 6.378212e-02 -7.114232e-02
#> Locus_2937 -3.177964e-02 3.052439e-02 2.774065e-02 4.791606e-02
#> Locus_2938 -9.808369e-03 -2.052324e-02 -6.629170e-03 5.990650e-02
#> Locus_2947 6.262756e-02 1.908629e-02 -4.009287e-02 -8.553416e-04
#> Locus_2949 3.538805e-02 -2.182309e-02 5.917409e-03 -1.256744e-02
#> Locus_2951 1.788055e-02 -3.667635e-02 -7.799690e-03 1.237235e-01
#> Locus_2953 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2954 -3.078872e-02 -8.674717e-06 -7.586947e-03 -2.712449e-02
#> Locus_2962 1.934702e-02 -1.020770e-02 -3.349364e-02 8.225682e-04
#> Locus_2966 5.098366e-02 5.935672e-02 2.593637e-02 1.058282e-01
#> Locus_2972 4.024449e-02 -2.655715e-03 -2.938672e-02 6.209118e-02
#> Locus_2974 -5.228103e-02 1.489615e-02 -2.470750e-03 3.931211e-02
#> Locus_2976 7.933021e-03 1.005313e-02 -1.360238e-02 6.886742e-02
#> Locus_2977 -2.448390e-02 -3.153851e-02 4.194346e-02 6.769173e-02
#> Locus_2978 -1.692329e-02 6.643077e-02 4.005450e-02 6.912488e-02
#> Locus_2983 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_2991 1.907729e-02 3.472171e-02 -1.598930e-02 -2.266532e-02
#> Locus_2993 1.361979e-02 -5.150633e-02 -1.641670e-02 3.738341e-02
#> Locus_2994 -7.422743e-03 3.010341e-02 2.463821e-02 -3.511463e-02
#> Locus_2995 -1.376766e-01 6.724705e-03 -5.798358e-02 -8.182830e-03
#> Locus_3005 -1.263016e-02 1.137431e-02 4.292948e-02 1.148079e-02
#> Locus_3009 -2.068099e-02 -1.047038e-02 -1.496784e-02 1.389263e-01
#> Locus_3010 5.866410e-02 9.338730e-03 -7.092032e-02 4.404352e-03
#> Locus_3012 8.168037e-02 -1.038133e-02 -3.104983e-02 1.684138e-01
#> Locus_3013 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3023 1.631561e-02 1.637245e-02 3.660683e-02 5.326551e-02
#> Locus_3029 1.499329e-02 -3.280054e-03 1.364774e-02 -1.911320e-02
#> Locus_3030 4.963426e-02 2.838331e-02 -4.747997e-02 2.768196e-02
#> Locus_3032 -8.125766e-03 -4.640478e-02 -2.565375e-02 6.527572e-02
#> Locus_3042 -2.585776e-02 2.032913e-02 2.202540e-02 9.612322e-03
#> Locus_3043 -7.443340e-02 1.210758e-02 1.885908e-02 2.030959e-02
#> Locus_3046 -3.266020e-02 -2.713894e-04 1.520647e-03 -2.788218e-02
#> Locus_3047 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3051 3.932845e-02 2.081057e-02 6.152783e-02 2.215438e-02
#> Locus_3054 -1.474439e-04 3.788992e-02 4.031903e-02 -3.860205e-02
#> Locus_3056 2.305332e-02 7.981472e-03 -3.930779e-02 -3.786126e-02
#> Locus_3058 -1.510782e-02 -5.358465e-02 -7.724908e-03 -4.894364e-02
#> Locus_3059 5.843623e-02 -3.529110e-02 -2.045923e-02 7.773991e-02
#> Locus_3060 -2.682731e-03 -9.369918e-03 2.111199e-03 9.201655e-03
#> Locus_3061 -1.206110e-02 -5.144052e-02 -5.456163e-03 -2.356595e-02
#> Locus_3062 8.434049e-02 7.694091e-02 1.524128e-02 7.247584e-02
#> Locus_3063 -1.063294e-01 4.529486e-02 1.735754e-02 2.279099e-02
#> Locus_3064 -1.028249e-01 4.966632e-03 -4.323366e-02 -1.130891e-02
#> Locus_3069 -4.250522e-02 -1.151759e-01 -2.083168e-02 -4.441345e-02
#> Locus_3078 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3079 -2.585776e-02 2.032913e-02 2.202540e-02 9.612322e-03
#> Locus_3081 2.443710e-02 -1.821699e-03 -7.522013e-03 3.766558e-03
#> Locus_3084 -2.818093e-02 -1.170768e-02 6.104915e-03 1.216318e-02
#> Locus_3087 -8.605558e-02 5.661564e-02 -4.890179e-02 9.761636e-03
#> Locus_3089 5.772383e-02 3.832559e-02 1.470905e-02 -4.885365e-03
#> Locus_3090 3.299924e-02 1.641819e-02 2.215046e-02 3.765163e-02
#> Locus_3091 7.187420e-02 5.667664e-02 -5.511376e-02 2.308590e-02
#> Locus_3092 4.976271e-02 1.519009e-04 7.025780e-04 -5.422066e-02
#> Locus_3093 3.414376e-02 -1.235613e-02 -8.223215e-04 2.621238e-03
#> Locus_3097 -5.004530e-03 2.896309e-03 -2.638515e-02 -1.440485e-02
#> Locus_3098 3.216090e-02 2.458318e-02 1.697926e-02 -9.794502e-03
#> Locus_3102 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3116 -3.078692e-02 2.102930e-02 -1.449004e-02 8.955419e-03
#> Locus_3122 1.507931e-02 1.112972e-01 4.618577e-02 4.255655e-02
#> Locus_3126 -2.486740e-02 1.968953e-02 8.549903e-02 2.548631e-02
#> Locus_3128 -3.234182e-02 -3.117681e-02 2.173775e-03 -5.699343e-02
#> Locus_3129 -6.491564e-03 6.064599e-03 2.024576e-02 1.197580e-02
#> Locus_3133 2.119859e-02 -2.452926e-02 -6.866149e-02 2.675156e-02
#> Locus_3138 2.528633e-02 4.517402e-02 3.492208e-02 9.151502e-02
#> Locus_3140 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3143 4.685723e-02 -3.215108e-02 -4.937649e-02 -2.517196e-01
#> Locus_3145 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3147 1.792885e-02 -8.717913e-02 1.960186e-02 -1.044242e-01
#> Locus_3148 -2.072124e-02 -3.619190e-02 -5.638986e-03 4.342636e-02
#> Locus_3154 -3.206060e-03 1.465060e-03 4.449782e-02 1.112903e-03
#> Locus_3155 -6.674996e-03 1.135608e-02 2.035686e-02 1.349176e-02
#> Locus_3160 -9.870788e-04 -4.786882e-02 2.948409e-03 5.021833e-02
#> Locus_3161 2.231254e-02 -4.717517e-02 8.331343e-02 -7.996694e-02
#> Locus_3164 -1.098213e-01 -3.911010e-02 -8.119569e-02 -2.380243e-02
#> Locus_3165 2.367336e-02 -2.035102e-02 -2.763265e-02 5.255573e-03
#> Locus_3167 6.058069e-03 -3.534700e-02 2.755408e-03 -4.634787e-02
#> Locus_3168 -8.343230e-03 2.334626e-02 1.550515e-02 3.019789e-02
#> Locus_3169 2.412412e-03 -5.715184e-02 4.032013e-03 2.338235e-02
#> Locus_3171 -3.666448e-03 -8.927997e-03 4.584859e-02 -1.976017e-02
#> Locus_3175 6.125108e-02 -3.707728e-02 7.844220e-03 1.931262e-02
#> Locus_3177 -2.800737e-02 -6.390008e-02 -3.574517e-03 4.471438e-03
#> Locus_3179 -1.179132e-02 -4.622001e-02 2.416104e-02 6.606886e-02
#> Locus_3181 1.633048e-02 -1.728696e-02 -1.351528e-02 -2.199359e-03
#> Locus_3182 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3188 1.061709e-01 1.404270e-01 5.923339e-02 1.003335e-01
#> Locus_3190 0.000000e+00 0.000000e+00 0.000000e+00 0.000000e+00
#> Locus_3191 1.555912e-02 3.230222e-02 -1.679538e-02 2.990276e-03
#> Locus_3192 -2.339503e-03 1.841427e-02 -1.667639e-02 2.658054e-02
#> Locus_3197 -1.234896e-01 -3.129068e-02 -3.926223e-02 6.232734e-02
#> Locus_3198 -1.613773e-03 -2.065823e-02 -1.211179e-02 4.170155e-02
#> PC2 PC3
#> Locus_10 -2.614296e-03 -1.546428e-03
#> Locus_15 -1.710141e-02 9.217899e-02
#> Locus_22 -1.477990e-02 -2.112234e-02
#> Locus_28 -2.088027e-02 9.904487e-03
#> Locus_32 -3.597126e-03 4.626218e-02
#> Locus_35 -8.935972e-18 -2.401430e-17
#> Locus_37 -3.299994e-17 1.721228e-17
#> Locus_61 -1.582734e-01 -3.192635e-02
#> Locus_71 -6.340163e-02 8.736774e-03
#> Locus_72 -2.579065e-16 1.204129e-16
#> Locus_82 -1.264313e-16 -2.544036e-18
#> Locus_84 -5.315106e-02 7.761020e-03
#> Locus_90 -1.899967e-02 1.835849e-01
#> Locus_91 -1.633687e-02 -7.872051e-02
#> Locus_96 -4.740268e-18 8.503443e-17
#> Locus_104 1.131104e-02 -1.452028e-03
#> Locus_122 -4.511764e-02 6.938763e-02
#> Locus_125 7.034268e-02 1.024283e-01
#> Locus_131 -3.418464e-02 -1.663661e-02
#> Locus_133 1.449511e-18 7.926980e-17
#> Locus_138 -8.034195e-02 -1.125334e-01
#> Locus_154 4.672107e-02 -7.171020e-02
#> Locus_162 1.559531e-01 1.324026e-01
#> Locus_166 -3.207078e-02 -5.474292e-02
#> Locus_177 -9.392174e-18 1.263897e-17
#> Locus_185 1.603569e-16 -4.577491e-17
#> Locus_191 1.642114e-02 2.302827e-02
#> Locus_193 1.983276e-02 -2.398045e-04
#> Locus_207 -8.534337e-02 -2.260005e-02
#> Locus_211 -7.092541e-03 -2.512722e-02
#> Locus_213 -3.122122e-02 -1.167299e-02
#> Locus_214 -2.040470e-02 -9.115165e-02
#> Locus_222 2.690249e-02 -7.340834e-03
#> Locus_227 5.375537e-02 4.419194e-02
#> Locus_242 3.524481e-02 7.603191e-02
#> Locus_249 -9.800163e-02 -7.456785e-02
#> Locus_254 3.938318e-02 1.458233e-03
#> Locus_256 -7.434068e-02 -1.310132e-01
#> Locus_263 -1.774134e-02 4.965224e-02
#> Locus_268 1.791371e-02 1.041619e-02
#> Locus_279 -7.788268e-03 9.786466e-03
#> Locus_285 4.400292e-02 3.714434e-02
#> Locus_287 -4.616942e-02 -1.138456e-02
#> Locus_292 -5.100330e-02 4.056116e-02
#> Locus_294 1.052136e-01 -6.090089e-02
#> Locus_303 -3.788172e-29 9.051762e-30
#> Locus_308 -8.803475e-03 1.510468e-02
#> Locus_312 1.058912e-01 -4.519878e-04
#> Locus_313 2.179103e-30 -9.148031e-31
#> Locus_314 4.845775e-02 6.392115e-02
#> Locus_317 -4.244805e-02 -4.791616e-03
#> Locus_318 4.693108e-02 -1.546672e-01
#> Locus_323 0.000000e+00 0.000000e+00
#> Locus_330 5.909861e-03 -1.280972e-02
#> Locus_331 2.113710e-04 7.462409e-03
#> Locus_333 -2.982422e-03 5.232459e-03
#> Locus_334 7.476153e-03 -2.959859e-02
#> Locus_336 -3.816487e-03 -1.558034e-02
#> Locus_337 3.483950e-03 9.060972e-02
#> Locus_339 -1.357122e-01 -1.607998e-01
#> Locus_341 3.360248e-02 2.193649e-01
#> Locus_345 -8.430844e-02 4.449696e-02
#> Locus_350 -2.475181e-03 8.258242e-02
#> Locus_354 1.791520e-02 3.533001e-02
#> Locus_359 -9.106179e-02 -1.788261e-01
#> Locus_365 -8.459447e-03 9.075843e-03
#> Locus_366 4.753655e-02 -2.779131e-02
#> Locus_369 -1.030199e-02 -9.456971e-03
#> Locus_372 -4.932711e-02 5.802349e-03
#> Locus_389 7.575067e-02 -5.612076e-02
#> Locus_391 -2.297922e-02 -5.730311e-02
#> Locus_392 -2.716454e-02 -9.869784e-02
#> Locus_393 -1.615607e-02 4.199535e-03
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#> Locus_1487 -2.921853e-02 -5.174408e-02
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#> Locus_1489 5.909861e-03 -1.280972e-02
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#> Locus_1694 -4.808372e-02 -5.138702e-02
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#> Locus_1984 3.868703e-02 -2.628365e-02
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#> Locus_1993 0.000000e+00 0.000000e+00
#> Locus_2003 4.226534e-03 -1.691598e-02
#> Locus_2007 2.757438e-02 -1.882851e-02
#> Locus_2014 5.035259e-02 8.774267e-02
#> Locus_2016 7.455677e-04 -4.595499e-03
#> Locus_2017 1.118459e-02 -4.154288e-02
#> Locus_2019 5.755248e-03 1.123407e-02
#> Locus_2020 9.424366e-03 4.146907e-03
#> Locus_2021 -3.467184e-02 1.218404e-02
#> Locus_2027 4.130319e-03 5.633112e-03
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#> Locus_2431 -9.823629e-03 -8.812916e-02
#> Locus_2432 6.267295e-02 -2.267247e-02
#> Locus_2433 -2.790245e-02 -3.537842e-02
#> Locus_2434 2.757438e-02 -1.882851e-02
#> Locus_2435 -1.858130e-02 -1.056683e-03
#> Locus_2436 -6.915570e-02 -4.334877e-02
#> Locus_2437 -4.479558e-02 -3.327405e-02
#> Locus_2439 1.448300e-02 -1.505714e-07
#> Locus_2448 -4.568670e-02 -7.295529e-02
#> Locus_2449 4.376680e-03 -4.297661e-02
#> Locus_2450 0.000000e+00 0.000000e+00
#> Locus_2451 -1.547197e-02 1.157009e-02
#> Locus_2452 0.000000e+00 0.000000e+00
#> Locus_2457 6.565398e-03 4.587187e-02
#> Locus_2461 -1.794457e-02 -2.573450e-02
#> Locus_2463 -3.744674e-02 -6.560037e-02
#> Locus_2466 0.000000e+00 0.000000e+00
#> Locus_2467 -8.636515e-02 -6.608745e-03
#> Locus_2473 0.000000e+00 0.000000e+00
#> Locus_2475 0.000000e+00 0.000000e+00
#> Locus_2477 -1.793283e-02 -6.013462e-02
#> Locus_2480 -1.214398e-02 -1.285462e-01
#> Locus_2494 0.000000e+00 0.000000e+00
#> Locus_2497 -2.592370e-02 -5.543555e-03
#> Locus_2501 -1.361493e-02 -1.906504e-02
#> Locus_2502 -1.561360e-02 1.968614e-03
#> Locus_2507 0.000000e+00 0.000000e+00
#> Locus_2510 -2.399572e-02 -8.753608e-02
#> Locus_2512 -1.223556e-01 1.515782e-02
#> Locus_2514 5.494332e-02 2.907319e-02
#> Locus_2517 -3.553494e-02 -1.398691e-01
#> Locus_2518 4.113785e-02 -2.591043e-02
#> Locus_2519 -3.904854e-02 5.542312e-02
#> Locus_2522 -7.567568e-02 -7.616425e-02
#> Locus_2526 -7.305174e-02 -5.358236e-02
#> Locus_2529 -1.630421e-02 2.936489e-02
#> Locus_2530 1.757171e-01 -8.201301e-02
#> Locus_2532 -3.686890e-02 -9.049864e-02
#> Locus_2540 -3.403153e-02 2.171328e-03
#> Locus_2542 5.908709e-02 -2.326681e-02
#> Locus_2546 0.000000e+00 0.000000e+00
#> Locus_2550 0.000000e+00 0.000000e+00
#> Locus_2555 -2.854963e-02 -1.150223e-02
#> Locus_2559 -1.157212e-01 6.298049e-02
#> Locus_2561 -2.656951e-02 7.896910e-02
#> Locus_2565 -1.403767e-02 -3.398986e-02
#> Locus_2566 2.757438e-02 -1.882851e-02
#> Locus_2567 9.818924e-03 1.545846e-02
#> Locus_2572 -6.029750e-02 -2.153872e-02
#> Locus_2573 5.750512e-02 -5.073335e-02
#> Locus_2577 -7.247875e-03 -3.053236e-02
#> Locus_2579 -2.719288e-02 6.425426e-02
#> Locus_2581 -2.209942e-02 -1.186604e-01
#> Locus_2583 8.481155e-02 -1.603403e-01
#> Locus_2588 -5.172440e-03 -6.681213e-02
#> Locus_2589 -4.408391e-02 1.308293e-01
#> Locus_2592 4.263537e-02 -2.435895e-02
#> Locus_2597 7.916051e-03 6.123541e-03
#> Locus_2598 -7.879184e-03 7.297317e-05
#> Locus_2599 -2.144760e-02 -2.346654e-02
#> Locus_2601 0.000000e+00 0.000000e+00
#> Locus_2602 -4.703170e-02 2.240656e-02
#> Locus_2611 -9.255436e-03 -1.299809e-02
#> Locus_2616 0.000000e+00 0.000000e+00
#> Locus_2617 -1.176026e-02 9.318362e-02
#> Locus_2619 -2.421850e-02 2.456830e-03
#> Locus_2620 0.000000e+00 0.000000e+00
#> Locus_2629 0.000000e+00 0.000000e+00
#> Locus_2632 -7.027450e-02 1.239378e-01
#> Locus_2634 -4.306282e-02 7.805629e-04
#> Locus_2636 -2.767303e-02 9.943019e-03
#> Locus_2639 -8.803475e-03 1.510468e-02
#> Locus_2644 1.096977e-01 -4.821636e-02
#> Locus_2647 6.796042e-02 -3.547761e-02
#> Locus_2648 -3.179272e-02 -2.626869e-03
#> Locus_2652 0.000000e+00 0.000000e+00
#> Locus_2655 -1.086571e-02 -3.253798e-02
#> Locus_2656 0.000000e+00 0.000000e+00
#> Locus_2657 -1.122416e-01 -2.916908e-03
#> Locus_2660 -3.273426e-02 1.796349e-02
#> Locus_2662 1.171436e-02 6.014013e-02
#> Locus_2665 -7.917429e-02 9.026693e-02
#> Locus_2668 -2.471800e-02 5.886071e-02
#> Locus_2669 9.307900e-02 -5.392080e-02
#> Locus_2670 4.797908e-02 -2.434321e-02
#> Locus_2671 -2.144760e-02 -2.346654e-02
#> Locus_2675 7.301871e-02 -1.900726e-02
#> Locus_2677 0.000000e+00 0.000000e+00
#> Locus_2685 -4.339616e-02 -1.337413e-01
#> Locus_2687 -2.076037e-03 -1.643702e-03
#> Locus_2696 -2.682440e-02 5.153771e-03
#> Locus_2698 -4.300490e-03 -1.139899e-01
#> Locus_2699 4.603944e-02 -2.527053e-02
#> Locus_2702 5.781562e-02 -3.189865e-02
#> Locus_2703 -8.305974e-02 -6.025512e-02
#> Locus_2705 -4.867665e-02 2.953401e-02
#> Locus_2712 -5.282369e-02 1.947705e-03
#> Locus_2718 1.134151e-02 1.190850e-02
#> Locus_2719 -6.688169e-02 1.609889e-02
#> Locus_2724 0.000000e+00 0.000000e+00
#> Locus_2725 -3.369047e-02 1.262875e-02
#> Locus_2727 0.000000e+00 0.000000e+00
#> Locus_2729 -2.088491e-02 5.565944e-02
#> Locus_2731 -7.660411e-03 -3.288079e-02
#> Locus_2733 -9.512830e-03 9.605248e-03
#> Locus_2736 -1.252751e-02 5.081093e-03
#> Locus_2738 1.104756e-02 4.861472e-03
#> Locus_2739 2.133917e-02 1.620115e-02
#> Locus_2753 -1.198239e-02 -1.524010e-02
#> Locus_2755 -2.573270e-02 1.732326e-02
#> Locus_2758 3.451701e-03 4.472838e-02
#> Locus_2760 6.530574e-02 -6.827507e-02
#> Locus_2761 6.267295e-02 -2.267247e-02
#> Locus_2764 1.104756e-02 4.861472e-03
#> Locus_2766 0.000000e+00 0.000000e+00
#> Locus_2767 9.286103e-03 -3.595227e-02
#> Locus_2768 -8.067946e-03 1.166107e-03
#> Locus_2770 5.071731e-02 -2.229829e-02
#> Locus_2774 2.757438e-02 -1.882851e-02
#> Locus_2776 9.571502e-02 4.721640e-02
#> Locus_2789 -9.641844e-02 -1.735078e-02
#> Locus_2792 3.911122e-02 1.682611e-02
#> Locus_2793 -1.631184e-02 1.553816e-02
#> Locus_2794 0.000000e+00 0.000000e+00
#> Locus_2795 -5.909551e-02 -7.220062e-02
#> Locus_2796 -7.032872e-03 1.561419e-02
#> Locus_2798 -8.803475e-03 1.510468e-02
#> Locus_2804 0.000000e+00 0.000000e+00
#> Locus_2806 0.000000e+00 0.000000e+00
#> Locus_2808 1.605847e-02 3.337292e-02
#> Locus_2809 1.846506e-02 -6.442017e-03
#> Locus_2810 -2.588149e-02 5.039872e-02
#> Locus_2811 -1.447530e-02 4.940214e-02
#> Locus_2814 2.698290e-03 8.076700e-03
#> Locus_2816 1.378719e-02 -9.414254e-03
#> Locus_2817 0.000000e+00 0.000000e+00
#> Locus_2818 -8.198791e-02 -3.923295e-02
#> Locus_2823 -9.035475e-02 -2.274580e-01
#> Locus_2830 -3.875340e-03 1.595488e-02
#> Locus_2831 -4.594043e-02 -2.975501e-02
#> Locus_2832 0.000000e+00 0.000000e+00
#> Locus_2835 -7.232578e-02 -3.189160e-02
#> Locus_2836 1.707294e-03 -1.314961e-02
#> Locus_2839 -1.353376e-01 -4.300697e-02
#> Locus_2842 6.464429e-02 5.561511e-02
#> Locus_2844 -1.327309e-02 1.221541e-02
#> Locus_2845 7.939836e-02 4.528258e-02
#> Locus_2850 4.108713e-02 -9.346073e-03
#> Locus_2852 -6.803008e-02 1.390371e-02
#> Locus_2854 -1.858130e-02 -1.056683e-03
#> Locus_2856 7.096855e-02 -1.718537e-02
#> Locus_2861 -3.322376e-02 6.675011e-02
#> Locus_2865 -5.150997e-03 -4.728486e-03
#> Locus_2868 0.000000e+00 0.000000e+00
#> Locus_2871 2.133917e-02 1.620115e-02
#> Locus_2872 0.000000e+00 0.000000e+00
#> Locus_2873 0.000000e+00 0.000000e+00
#> Locus_2874 -8.744685e-02 5.347873e-03
#> Locus_2875 5.909861e-03 -1.280972e-02
#> Locus_2878 4.723265e-02 1.382078e-02
#> Locus_2881 -5.549037e-03 5.456633e-03
#> Locus_2884 -1.636894e-02 -7.603246e-03
#> Locus_2890 1.714212e-02 -8.009065e-03
#> Locus_2895 0.000000e+00 0.000000e+00
#> Locus_2896 3.972971e-02 -1.403280e-02
#> Locus_2904 -9.396840e-02 -3.392391e-02
#> Locus_2905 0.000000e+00 0.000000e+00
#> Locus_2910 -5.968260e-03 1.430441e-02
#> Locus_2915 -3.716147e-02 -1.889408e-01
#> Locus_2916 -1.926399e-02 -7.350915e-02
#> Locus_2923 -1.421650e-02 -2.864848e-02
#> Locus_2932 -5.269974e-02 1.439119e-02
#> Locus_2933 -4.510642e-03 -3.717145e-03
#> Locus_2934 -5.892366e-02 -4.342821e-02
#> Locus_2936 6.270243e-03 -7.395411e-02
#> Locus_2937 -1.633048e-01 -1.417669e-02
#> Locus_2938 -1.739276e-02 -2.325277e-02
#> Locus_2947 -1.020096e-01 -5.282935e-02
#> Locus_2949 -4.235331e-02 1.747914e-02
#> Locus_2951 -4.382679e-02 -4.147274e-02
#> Locus_2953 0.000000e+00 0.000000e+00
#> Locus_2954 -1.176026e-02 9.318362e-02
#> Locus_2962 -4.716795e-03 -2.928802e-02
#> Locus_2966 -3.356713e-02 5.354542e-02
#> Locus_2972 -5.178229e-02 -2.173316e-01
#> Locus_2974 -5.716469e-02 -6.187819e-03
#> Locus_2976 -1.605264e-02 1.777930e-02
#> Locus_2977 1.381067e-02 -5.219291e-02
#> Locus_2978 -1.902606e-02 -1.700275e-02
#> Locus_2983 0.000000e+00 0.000000e+00
#> Locus_2991 4.964319e-03 7.679423e-03
#> Locus_2993 -6.855761e-02 -4.290475e-02
#> Locus_2994 1.939634e-03 9.273143e-04
#> Locus_2995 1.253459e-01 -4.534494e-02
#> Locus_3005 -7.447970e-03 -2.070206e-02
#> Locus_3009 1.117201e-01 -1.016689e-01
#> Locus_3010 -1.324281e-01 7.400401e-02
#> Locus_3012 8.921171e-02 1.447456e-01
#> Locus_3013 0.000000e+00 0.000000e+00
#> Locus_3023 -1.689622e-02 -8.877949e-02
#> Locus_3029 -3.978531e-02 1.564716e-02
#> Locus_3030 -2.676695e-02 -1.168646e-01
#> Locus_3032 -6.029750e-02 -2.153872e-02
#> Locus_3042 -3.233519e-02 -3.959477e-04
#> Locus_3043 1.834557e-02 -4.373100e-02
#> Locus_3046 5.544263e-02 -9.430631e-03
#> Locus_3047 0.000000e+00 0.000000e+00
#> Locus_3051 9.739818e-02 -7.494598e-02
#> Locus_3054 -9.821209e-03 4.459011e-02
#> Locus_3056 -1.519880e-03 9.233913e-02
#> Locus_3058 -5.581135e-02 -2.475936e-01
#> Locus_3059 -3.434094e-02 5.947456e-02
#> Locus_3060 -8.756190e-03 -2.855573e-02
#> Locus_3061 -1.842140e-02 3.393064e-03
#> Locus_3062 9.145338e-02 -8.809134e-03
#> Locus_3063 1.821985e-01 -1.326319e-01
#> Locus_3064 9.792471e-02 -3.322853e-02
#> Locus_3069 -8.884651e-02 -7.193522e-02
#> Locus_3078 0.000000e+00 0.000000e+00
#> Locus_3079 -3.233519e-02 -3.959477e-04
#> Locus_3081 5.909861e-03 -1.280972e-02
#> Locus_3084 7.984753e-02 -4.381937e-02
#> Locus_3087 2.703336e-02 2.893306e-02
#> Locus_3089 3.241095e-03 2.033642e-02
#> Locus_3090 2.544017e-02 -1.686437e-02
#> Locus_3091 -3.440993e-02 -2.246096e-02
#> Locus_3092 -1.711819e-02 -3.928254e-02
#> Locus_3093 -1.136339e-02 -4.562299e-03
#> Locus_3097 2.113710e-04 7.462409e-03
#> Locus_3098 -1.711493e-02 -1.492050e-03
#> Locus_3102 0.000000e+00 0.000000e+00
#> Locus_3116 3.225225e-02 -1.585627e-02
#> Locus_3122 -1.028494e-01 8.100550e-02
#> Locus_3126 -1.430819e-02 -5.220864e-02
#> Locus_3128 1.165995e-01 -3.641667e-02
#> Locus_3129 -2.468538e-03 -2.695415e-02
#> Locus_3133 1.193652e-02 -2.860882e-02
#> Locus_3138 4.059503e-02 -3.252279e-03
#> Locus_3140 0.000000e+00 0.000000e+00
#> Locus_3143 -4.823805e-02 9.809943e-03
#> Locus_3145 0.000000e+00 0.000000e+00
#> Locus_3147 -5.250548e-02 -1.213548e-01
#> Locus_3148 -3.970111e-03 -2.212356e-02
#> Locus_3154 -2.332279e-02 2.158191e-02
#> Locus_3155 -5.640503e-03 -2.840603e-02
#> Locus_3160 -9.808816e-02 -1.608655e-02
#> Locus_3161 -3.169027e-02 -5.257177e-02
#> Locus_3164 2.437094e-02 1.078840e-02
#> Locus_3165 -3.158070e-03 8.297162e-03
#> Locus_3167 -1.660613e-02 2.900666e-03
#> Locus_3168 -6.522472e-03 -1.137442e-02
#> Locus_3169 -6.596724e-02 -3.861774e-02
#> Locus_3171 2.202652e-02 -5.687919e-02
#> Locus_3175 -4.427675e-02 2.358290e-02
#> Locus_3177 -8.264310e-02 -4.782030e-02
#> Locus_3179 -6.059097e-02 -1.273264e-02
#> Locus_3181 -2.052152e-02 -2.973971e-02
#> Locus_3182 0.000000e+00 0.000000e+00
#> Locus_3188 -4.648820e-02 -1.773875e-02
#> Locus_3190 0.000000e+00 0.000000e+00
#> Locus_3191 -6.869141e-03 -1.780904e-02
#> Locus_3192 5.907364e-02 3.470630e-02
#> Locus_3197 1.560651e-01 -1.020570e-01
#> Locus_3198 -2.825616e-02 -2.030831e-02
#>
#> $sites
#> RDA1 RDA2 RDA3 PC1 PC2 PC3
#> ALT3 -3.04652568 0.18212478 -1.03792529 -0.18125460 1.19051626 -0.81291564
#> BAR360 1.30674241 0.16343426 -0.80667863 -0.88739574 -0.67411286 0.08499438
#> BLL5 -0.49561740 0.28265102 -1.81340536 -1.24385121 0.01825177 0.64437486
#> BNT5 1.40006492 2.26680990 -0.90363212 -1.60173649 0.47697542 0.20989270
#> BOF1 0.02237711 -0.58659179 0.35290055 0.09616041 0.13024220 -0.17068122
#> BOT3 1.17354392 -0.09846946 -1.38996703 1.99555088 -0.56848391 0.21692739
#> CAR1 0.13944774 0.11401119 0.54448661 -1.17305094 1.12532884 0.56242122
#> CTR7 0.57463960 -1.41913880 -0.76843654 0.69748227 -1.09790882 -2.65300190
#> CYR26 -0.17679198 1.82968703 -0.60920177 -0.78408649 -0.69666262 0.10069270
#> FCT13 -1.50065311 0.58236200 0.94955915 0.78892038 -2.07117041 -0.31182302
#> HAY2 1.09681816 2.23624452 -0.82369942 3.31624927 -0.65566763 0.75153843
#> HHR2 0.40392939 -0.26899900 -0.56629157 0.69839742 -0.88633433 -2.29509006
#> HNT18 0.57951586 0.37914426 0.66380981 -0.34802980 -0.90307087 0.84874839
#> HOP1 0.99949363 -3.61093727 1.92983022 -3.27890361 -1.21214499 -2.93500514
#> INK1 -0.62431770 -2.43530133 1.52904501 1.64167105 1.72613338 1.26809826
#> INK6 -0.76144069 -1.94768029 1.45620471 0.72752273 1.31089236 3.15421619
#> KBG6 -1.12316366 1.29902872 1.76919724 -0.89576522 -0.57192111 1.67540153
#> KRV3 -0.83011567 1.03241182 2.92337626 -1.15499107 -0.51535596 1.23517790
#> LBH40 -3.38343934 0.78021197 -1.16206130 -0.27783572 -2.88780145 0.13704048
#> LCV19 1.41730659 0.59371202 -0.43494475 -0.99395214 -0.44478574 -0.40830211
#> LFV10 2.17801418 1.58398090 -2.85752101 -0.19428704 0.05461702 -0.87864713
#> LTF1 -1.19028780 1.68429380 -0.35290837 0.95455005 1.59444765 -0.55626456
#> LTF6 -0.20401877 2.31565650 -0.18181732 -0.81192683 0.97670179 -0.12538182
#> MLG2 1.26242424 -1.33186662 -0.68815927 0.13179948 1.25105297 0.39990416
#> MLG5 2.24522263 -1.23972506 -1.62344668 -1.54173507 0.29578629 0.49952795
#> MNR2 1.34408027 3.12640133 1.21820589 1.23278104 -0.38008758 0.65214050
#> MRO3 -0.90677205 -0.77060029 -0.71564860 -0.72320959 -0.22178538 3.18547602
#> OLA14 0.15255857 -1.11919417 0.56675740 -1.18512783 -1.60448511 -0.09124397
#> OLA9 0.28094559 -2.19056697 0.49395406 2.18272646 -0.32955156 -1.34535403
#> POT1 1.19467655 1.07827084 -3.21337129 -0.90679760 -0.10609221 0.91504443
#> PRU1 -1.34146285 0.16023842 1.31252209 -1.78438402 1.74347044 0.09718033
#> PRU6 -1.34137160 -0.06955285 1.84357022 -0.09405434 3.35866648 -2.10625633
#> PT8 2.68122107 1.92407405 3.05931060 2.03856146 1.01231838 -0.26512237
#> REY5 0.01458264 0.35146050 0.16566456 -0.80735633 1.68861598 0.72646257
#> RPS38 -0.10736616 -1.40303764 -0.66948119 1.01780814 -1.38337579 -0.05312248
#> RPS9 0.36300899 -1.28702124 -0.38240402 1.80045288 -1.21995186 -0.87680557
#> SBR10 -1.07199222 -0.19265149 -1.23792726 -0.72244158 0.68354660 0.52876434
#> SKY3 1.84547049 0.01989479 -0.82772302 2.63041386 -0.73046854 0.78369396
#> SKY5 1.55427227 -0.71011289 -1.98228379 0.72598494 -0.75219186 1.78410360
#> SRM1 0.84534245 -0.91395100 0.31451775 1.21263595 1.18442610 -1.19110629
#> SRM7 1.04011716 0.23696799 0.21663185 0.44339227 -1.70176267 0.53311930
#> SUT7 -0.78322966 0.06093197 2.09272163 -2.22020194 -0.80921546 0.20545489
#> TTC1 -0.02128617 -0.12341451 -0.02633965 -0.13090227 0.27389739 0.12536882
#> TTC2 -0.17932955 -0.06541184 0.37291947 0.21800993 0.05075211 -0.93296482
#> WAW1 -0.78599742 -0.90595824 -0.04550280 -0.32804932 0.26131127 2.31931165
#> WDC1 -0.24559603 0.78823272 1.51112802 -0.17364688 -0.15607373 0.66523351
#> WDC2 0.27725291 0.27007528 1.63824140 1.16500631 -0.48705431 -2.45284480
#> WHR2 -0.07914718 -3.67615862 -0.79373081 -1.70685774 -1.85198852 -2.02632247
#> WTW14 0.98422269 -0.99308833 0.07526736 -0.09889776 -1.49153584 0.71215996
#> WTW3 0.76091603 -0.96677881 0.43809916 0.84684508 1.31227123 -0.85409273
#> YUC3 0.12124716 1.93676658 1.78527862 0.04109831 -0.72095501 0.27763316
#> ZUM18 -4.36991029 0.76779695 -1.95737024 -0.62322698 3.04396981 -0.91151007
#> ZUM20 -3.68962221 0.27933242 -1.35132055 0.26993556 2.36780441 -1.04624505
#>
#> $constraints
#> RDA1 RDA2 RDA3 PC1 PC2
#> ALT3 -2.30232603 0.178476566 -1.273124800 -0.18125460 1.19051626
#> BAR360 1.37940868 -0.065126700 -0.646064312 -0.88739574 -0.67411286
#> BLL5 -0.43213837 0.250094636 -2.278343132 -1.24385121 0.01825177
#> BNT5 1.44454669 1.471689545 -0.532155498 -1.60173649 0.47697542
#> BOF1 -0.06741513 -1.159963268 0.174289626 0.09616041 0.13024220
#> BOT3 0.55754345 -0.975193831 -1.537953693 1.99555088 -0.56848391
#> CAR1 1.79891628 -0.389509726 0.266500487 -1.17305094 1.12532884
#> CTR7 0.03071892 -1.427592554 -0.520972326 0.69748227 -1.09790882
#> CYR26 -0.15160265 3.387710248 -1.647167917 -0.78408649 -0.69666262
#> FCT13 -1.80972775 0.414120411 0.777307321 0.78892038 -2.07117041
#> HAY2 -0.58060672 1.567252812 -0.722616064 3.31624927 -0.65566763
#> HHR2 -0.16423724 0.350876087 0.008011260 0.69839742 -0.88633433
#> HNT18 -0.29940023 0.302425471 1.052818842 -0.34802980 -0.90307087
#> HOP1 1.47756297 -2.038349074 1.647901232 -3.27890361 -1.21214499
#> INK1 -0.60465735 -2.202837767 0.923984182 1.64167105 1.72613338
#> INK6 -0.51019862 -2.347261234 0.994390300 0.72752273 1.31089236
#> KBG6 -0.74968503 2.026939503 2.164817885 -0.89576522 -0.57192111
#> KRV3 -0.91524327 1.059044044 2.964440647 -1.15499107 -0.51535596
#> LBH40 -3.70358745 0.935639828 -1.086405521 -0.27783572 -2.88780145
#> LCV19 0.94811302 0.582489237 -0.859764314 -0.99395214 -0.44478574
#> LFV10 2.48093130 1.405579757 -2.656816123 -0.19428704 0.05461702
#> LTF1 -0.35610701 1.637391587 0.021917909 0.95455005 1.59444765
#> LTF6 -0.04964522 1.971124227 0.147805504 -0.81192683 0.97670179
#> MLG2 1.24503917 -1.199453777 -0.877007640 0.13179948 1.25105297
#> MLG5 1.48663836 -0.782105364 -0.848109947 -1.54173507 0.29578629
#> MNR2 1.23002491 2.540925090 1.922764606 1.23278104 -0.38008758
#> MRO3 -0.95445209 -1.013844280 -1.409973257 -0.72320959 -0.22178538
#> OLA14 -0.30079134 -1.016598326 0.321300347 -1.18512783 -1.60448511
#> OLA9 0.07146681 -1.186361181 0.574998137 2.18272646 -0.32955156
#> POT1 1.91990376 1.808688216 -3.208075117 -0.90679760 -0.10609221
#> PRU1 0.11453380 -0.161962958 1.361084604 -1.78438402 1.74347044
#> PRU6 0.20159550 0.065433696 1.347503956 -0.09405434 3.35866648
#> PT8 2.11874422 1.509877281 1.916136965 2.03856146 1.01231838
#> REY5 1.05140964 -0.651575539 -0.197802957 -0.80735633 1.68861598
#> RPS38 -0.28115349 -1.111600218 -0.584670599 1.01780814 -1.38337579
#> RPS9 -0.06967421 -0.891913458 -0.522923014 1.80045288 -1.21995186
#> SBR10 -1.92148578 -0.091958527 -1.118955802 -0.72244158 0.68354660
#> SKY3 1.26561864 -0.699170450 -0.451941923 2.63041386 -0.73046854
#> SKY5 1.25734916 -0.824512883 -0.431379854 0.72598494 -0.75219186
#> SRM1 0.73072164 -0.092175984 -0.003457347 1.21263595 1.18442610
#> SRM7 0.75160889 0.123358532 -0.055903780 0.44339227 -1.70176267
#> SUT7 -0.59130451 0.628288883 1.979690022 -2.22020194 -0.80921546
#> TTC1 0.01583928 -0.456916219 -0.009593412 -0.13090227 0.27389739
#> TTC2 0.03392901 -0.264150708 -0.031079054 0.21800993 0.05075211
#> WAW1 -0.51603287 -1.897578499 0.341418542 -0.32804932 0.26131127
#> WDC1 -0.51422518 0.001574274 1.949132991 -0.17364688 -0.15607373
#> WDC2 -0.57638222 0.980590933 1.757803316 1.16500631 -0.48705431
#> WHR2 -0.52543655 -2.544281462 -0.812555186 -1.70685774 -1.85198852
#> WTW14 0.83816479 -0.909642385 0.578514651 -0.09889776 -1.49153584
#> WTW3 0.86440999 -0.429580688 0.509129960 0.84684508 1.31227123
#> YUC3 -0.42307576 1.341288928 1.124574021 0.04109831 -0.72095501
#> ZUM18 -2.93472429 0.138528634 -1.229777594 -0.62322698 3.04396981
#> ZUM20 -3.00942250 0.151808636 -1.273647132 0.26993556 2.36780441
#> PC3
#> ALT3 -0.81291564
#> BAR360 0.08499438
#> BLL5 0.64437486
#> BNT5 0.20989270
#> BOF1 -0.17068122
#> BOT3 0.21692739
#> CAR1 0.56242122
#> CTR7 -2.65300190
#> CYR26 0.10069270
#> FCT13 -0.31182302
#> HAY2 0.75153843
#> HHR2 -2.29509006
#> HNT18 0.84874839
#> HOP1 -2.93500514
#> INK1 1.26809826
#> INK6 3.15421619
#> KBG6 1.67540153
#> KRV3 1.23517790
#> LBH40 0.13704048
#> LCV19 -0.40830211
#> LFV10 -0.87864713
#> LTF1 -0.55626456
#> LTF6 -0.12538182
#> MLG2 0.39990416
#> MLG5 0.49952795
#> MNR2 0.65214050
#> MRO3 3.18547602
#> OLA14 -0.09124397
#> OLA9 -1.34535403
#> POT1 0.91504443
#> PRU1 0.09718033
#> PRU6 -2.10625633
#> PT8 -0.26512237
#> REY5 0.72646257
#> RPS38 -0.05312248
#> RPS9 -0.87680557
#> SBR10 0.52876434
#> SKY3 0.78369396
#> SKY5 1.78410360
#> SRM1 -1.19110629
#> SRM7 0.53311930
#> SUT7 0.20545489
#> TTC1 0.12536882
#> TTC2 -0.93296482
#> WAW1 2.31931165
#> WDC1 0.66523351
#> WDC2 -2.45284480
#> WHR2 -2.02632247
#> WTW14 0.71215996
#> WTW3 -0.85409273
#> YUC3 0.27763316
#> ZUM18 -0.91151007
#> ZUM20 -1.04624505
#>
#> $biplot
#> RDA1 RDA2 RDA3 PC1 PC2 PC3
#> CA_rPCA1 -0.3708764 0.31873717 0.7861656 0 0 0
#> CA_rPCA2 -0.3372560 -0.09456548 -0.0628678 0 0 0
#> CA_rPCA3 0.2301084 -0.44498596 0.5823455 0 0 0
#>
#> $call
#> rda(formula = gen ~ CA_rPCA1 + CA_rPCA2 + CA_rPCA3 + Condition(PC1 +
#> PC2 + PC3 + x + y), data = moddf)
#>
#> $tot.chi
#> [1] 143.389Variance partitioning with partial RDA
In many cases, we want to understand the independent contributions of each explanatory variable in combination with our covariables. Also, we can understand how much confounded variance there is (i.e., variation that is explained by the combination of multiple explanatory variables). The best way to do this is to run partial RDAs on each set of explanatory variables and examine the inertia of each pRDA result as it compares to the full model. In this case, the full model is an RDA that considers neutral processes as explanatory variables rather than conditioning variables (covariables).
The call for this full model looks like this:
rda(gen ~ env + covariable, data), rather than a true
partial RDA which would take the form:
rda(gen ~ env + Condition(covariable), data).
In algatr, we can run variance partitioning using the function
rda_varpart(), which will run the full model (as above) and
then will run successive pRDAs which are conditioned on three successive
sets of variables. These variable sets are significant environmental
variables (in our case, environmental PCs 2 and 3), population genetic
structure (once again determined using a PC with a user-specified number
of PC axes), and geography (sampling coordinates). This function will
then provide us with information on how variance is partitioned in the
form of a table containing relevant statistics; this table is nearly
identical to Table 2 from Capblancq
& Forester 2021. The rda_varpart() parameters are
identical to those from rda_run(). As with
rda_run(), we can provide a number for the number of PCs to
consider for population structure (nPC), or this argument
can be set to "manual" if a user would like to manually
enter a number into the console after viewing the screeplot. We can then
use rda_varpart_table() to visualize these results in an
aesthetically pleasing way; users can specify whether to display the
function call in the table (call_col; defaults to
FALSE) and the number of digits to round to
digits (the default is 2).
varpart <- rda_varpart(gen, env, liz_coords,
Pin = 0.05, R2permutations = 1000,
R2scope = T, nPC = 3
)
#> Step: R2.adj= 0
#> Call: gen ~ 1
#>
#> R2.adjusted
#> <All variables> 0.0424326339
#> + CA_rPCA2 0.0196214014
#> + CA_rPCA3 0.0137427906
#> + CA_rPCA1 0.0009953656
#> <none> 0.0000000000
#>
#> Df AIC F Pr(>F)
#> + CA_rPCA2 1 264.09 2.0407 0.004 **
#> ---
#> Signif. codes: 0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
#>
#> Step: R2.adj= 0.0196214
#> Call: gen ~ CA_rPCA2
#>
#> R2.adjusted
#> <All variables> 0.04243263
#> + CA_rPCA3 0.03919420
#> + CA_rPCA1 0.01991617
#> <none> 0.01962140
#>
#> Df AIC F Pr(>F)
#> + CA_rPCA3 1 263.97 2.0389 0.006 **
#> ---
#> Signif. codes: 0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
#>
#> Step: R2.adj= 0.0391942
#> Call: gen ~ CA_rPCA2 + CA_rPCA3
#>
#> R2.adjusted
#> <All variables> 0.04243263
#> + CA_rPCA1 0.04243263
#> <none> 0.03919420
#>
#> Df AIC F Pr(>F)
#> + CA_rPCA1 1 264.72 1.1691 0.19
rda_varpart_table(varpart, call_col = TRUE)| Variance partitioning | |||||||
| Model | pRDA model call | Inertia | R2 | Adjusted R2 | p (>F) | Prop. of explainable variance | Prop. of total variance |
|---|---|---|---|---|---|---|---|
| Full model | gen ~ CA_rPCA2 + CA_rPCA3 + PC1 + PC2 + PC3 + x + y | 52.67 | 0.37 | 0.27 | 0.00 | 1.00 | 0.37 |
| Pure enviro. model | gen ~ CA_rPCA2 + CA_rPCA3 + Condition(PC1 + PC2 + PC3 + x + y) | 4.61 | 0.03 | 0.00 | 0.06 | 0.09 | 0.03 |
| Pure pop. structure model | gen ~ PC1 + PC2 + PC3 + Condition(CA_rPCA2 + CA_rPCA3 + x + y) | 23.09 | 0.16 | 0.13 | 0.00 | 0.44 | 0.16 |
| Pure geography model | gen ~ x + y + Condition(PC1 + PC2 + PC3 + CA_rPCA2 + CA_rPCA3) | 5.32 | 0.04 | 0.01 | 0.00 | 0.10 | 0.04 |
| Confounded variance | 19.66 | 0.37 | 0.14 | ||||
| Total unexplained variance | 90.72 | 0.63 | |||||
| Total inertia | 143.39 | 1.00 | |||||
From the above table, we can see that the full model only explains 27% of the total variance in our genetic data. Furthermore, our pure environmental model explains only a small proportion of the variance (“inertia”) contained within the full model (9%), and that population structure explains substantially more (44%). There is also a large amount of variance (37%) within the full model that is confounded, meaning that variables within the three variable sets explain a substantial amount of the full model’s variance when in combination with one another. This is a relatively common occurrence (see Capblancq & Forester 2021), and users should always keep in mind how collinearity may affect their results (and account for this, if possible; see algatr’s environmental data vignette for more information on how to do so). However, accounting for collinearity may also elevate false positive rates, while failing to account for it at all can lead to elevated false negative rates.
In our case, it is clear from our variance partitioning analysis that much of the variation in our genetic data can explained by population structure, and thus users may want to further explore more sophisticated approaches for quantifying demographic history or population structure before moving forward with outlier loci detection. As with most analyses, careful consideration of the input data and the biology of the system are critical to draw robust and reliable conclusions.
Identifying candidate SNPs with rda_getoutliers()
Having run variance partitioning on our data, we can now move forward with detecting outlier loci that are significantly associated with environmental variables. To do so, we’ll first run a partial RDA on only those significant environmental variables, and we’ll also want to add population structure as a covariable (i.e., a conditioning variable). We’ll only retain two PC axes for our measurement of population structure.
mod_pRDA <- rda_run(gen, env, model = "best", correctPC = TRUE, nPC = 2)
#> Step: R2.adj= 0
#> Call: gen ~ 1
#>
#> R2.adjusted
#> <All variables> 0.0202527037
#> + CA_rPCA2 0.0196214014
#> + CA_rPCA3 0.0137427906
#> + CA_rPCA1 0.0009953656
#> <none> 0.0000000000
#> + Condition(PC1 + PC2) 0.0000000000
#>
#> Df AIC F Pr(>F)
#> + CA_rPCA2 1 264.09 2.0407 0.004 **
#> ---
#> Signif. codes: 0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1
#>
#> Step: R2.adj= 0.0196214
#> Call: gen ~ CA_rPCA2
#>
#> R2.adjusted
#> + CA_rPCA3 0.03919420
#> <All variables> 0.02025270
#> + CA_rPCA1 0.01991617
#> <none> 0.01962140
#> + Condition(PC1 + PC2) 0.01834925Given this partial RDA, only environmental PC2 is considered significant.
mod_pRDA$anova
#> R2.adj Df AIC F Pr(>F)
#> + CA_rPCA2 0.019621 1 264.09 2.0407 0.004 **
#> <All variables> 0.020253
#> ---
#> Signif. codes: 0 '***' 0.001 '**' 0.01 '*' 0.05 '.' 0.1 ' ' 1Now that we have the RDA model, we can discover SNPs that are
outliers. In algatr, this can be done using two different methods to
detect outlier loci, specified by the outlier_method
argument. The first uses Z-scores for outlier detection, and is from Forester
et al. 2018; their code can be found here. The
other method transforms RDA loadings into p-values, and was developed by
Capblancq
et al. 2018 and is further described in Capblancq
& Forester 2021; code developed to run this is associated with
Capblancq and Forester 2021 and can be found here. We
provide brief explanations for each of these methods below.
N.B.: Some researchers may prefer to do MAF filtering and linkage disequilibrium pruning during outlier detection (e.g., Capblancq & Forester 2021). In this case, only one SNP per contig is selected based on the highest loading. The algatr pipeline does this filtering initially when processing genetic data; see data processing vignette for more information.
Identifying outliers using the Z-scores method
To understand how the Z-scores method works to detect outliers, let’s
first take a look at the loadings of SNPs on the first two RDA axes
using the rda_plot() function. This will show us histograms
for each of the selected RDA axes. SNPs that are not associated with
environmental variables will fall in the center of these distributions,
with outliers falling at the edges. The Z-score method works by setting
a Z-score (number of standard deviations) threshold beyond which SNPs
will be considered outliers (see here for
further details).
If we set this value too stringently, we minimize our false positive rate but risk detecting false negatives; if it’s set too high, we increase our false positive rate. Researchers will want to adjust this parameter based on their tolerance for detecting false positives, but also the strength of selection under which outliers are subject to (assuming that SNPs that are more significantly associated with environmental variables are also under stronger selection).
Because only one environmental variable was associated with our genetic data, we only have one resulting RDA axis.
rda_plot(mod_pRDA, axes = "all", binwidth = 20)
Now, let’s discover outliers with the rda_getoutliers()
function. The method is specified using the outlier_method
argument, and the number of standard deviations to detect outliers is
specified by the z argument; it defaults to 3. The
resulting object is a dataframe containing the SNP name, the RDA axis
it’s associated with, and the loading along this axis. A screeplot for
the RDA axes is automatically generated (but can be switched off using
the plot argument).
rda_sig_z <- rda_getoutliers(mod_pRDA, naxes = "all", outlier_method = "z", z = 3, plot = FALSE)
# How many outlier SNPs were detected?
length(rda_sig_z$rda_snps)
#> [1] 19Identifying outliers using the p-value method
Another method to detect outliers is a p-value method wherein rather
than using the loadings along RDA axes to identify extreme outliers (as
with the Z-scores method above), the loadings are transformed into
p-values. To do so, Mahalanobis distances are calculated for each locus,
scaled based on a genomic inflation factor, from which p-values and
q-values can be computed. We then adjust our p-values (based on a false
discovery rate, for example; this adjustment is done using the
p.adjust() function) and subsequently identify outliers
based on a user-specified significance threshold. Rather than detecting
outliers based on a p-value significance threshold, one could also
detect outliers based on q-values (which are also provided as output in
algatr), as is done in Capblancq
et al. 2018 and Capblancq
& Forester 2021. To do so, we would select loci that fall below
a user-specified threshold based on the proportion of false positives we
allow. For example, if we allow 10% or fewer outliers to be false
positives, outliers could be identified based on loci that have q-values
=< 0.1.
Now, let’s detect outliers using this method. To do so, along with
outlier_method, we need to specify p_adj,
which is the method by which p-values are adjusted, as well as
sig, which is our significance threshold. We will also tell
the function not to produce a screeplot.
One limitation of the p-value outlier method is that you cannot
calculate outliers given fewer than two RDA axes (as is the case with
our partial RDA results). So, let’s move forward with this outlier
detection method but using the results from our simple RDA model with
variable selection, mod_best.
rda_sig_p <- rda_getoutliers(mod_best, naxes = "all", outlier_method = "p", p_adj = "fdr", sig = 0.01, plot = FALSE)
# How many outlier SNPs were detected?
length(rda_sig_p$rda_snps)
#> [1] 101As mentioned above, we could also identify outliers based on
q-values. To do so, we can extract the q-values from the resulting
object from rda_getoutliers().
# Identify outliers that have q-values < 0.1
q_sig <-
rda_sig_p$rdadapt %>%
# Make SNP names column from row names
mutate(snp_names = row.names(.)) %>%
filter(q.values <= 0.1)
# How many outlier SNPs were detected?
nrow(q_sig)
#> [1] 170Note the differences between the two methods: using the p-value method, we found many more outlier SNPs than with the Z-scores method (101 vs. 19). Even more outliers were found when only using q-values as a significant threshold (170). We can also see that 19 SNPs were detected using all three methods.
Reduce(intersect, list(
q_sig$snp_names,
rda_sig_p$rda_snps,
rda_sig_z$rda_snps
))
#> [1] "Locus_517" "Locus_771" "Locus_786" "Locus_945" "Locus_1247"
#> [6] "Locus_1263" "Locus_1278" "Locus_1318" "Locus_1374" "Locus_1439"
#> [11] "Locus_1524" "Locus_1801" "Locus_1949" "Locus_2045" "Locus_2338"
#> [16] "Locus_2431" "Locus_2559" "Locus_2665" "Locus_2947"Visualizing RDA results with rda_plot()
algatr provides options for visualizing RDA results in two ways with
the rda_plot() function: a biplot and a Manhattan plot.
RDA biplot
Let’s build a biplot (or some call it a triplot, since it also shows
loadings of environmental variables) with RDA axes 1 and 2, specified
using the rdaplot argument. In this case, we also need to
specify the number of axes to display using the biplot_axes
argument. The default of rda_plot() is to plot all
variables, regardless of whether they were considered significant or
not.
# In the case of our partial RDA, there was only one RDA axis, so a histogram is generated
rda_plot(mod_pRDA, rda_sig_z$rda_snps, rdaplot = TRUE, manhattan = FALSE, binwidth = 0.01)
Manhattan plot
Let’s now build a Manhattan plot with a significance level of 0.05
using the manhattan argument; if this is set to TRUE, we
must also provide a significance threshold value using the
sig argument (the default is 0.05). Be aware that outliers
were detected with a different p-value threshold which is why there are
grey points above our threshold line (i.e., data points are colorized
according to the threshold of the model, not the visualization).
rda_plot(mod_best, rda_sig_p$rda_snps, rda_sig_p$pvalues, rdaplot = FALSE, manhattan = TRUE)
Interpreting RDA results
As discussed in Capblancq & Forester 2021, users should be aware that “statistical selection of variables optimizes variance explained but cannot, by itself, identify the ecological or mechanistic drivers of genetic variation.”
Identifying environmental associations with rda_cor()
and rda_table()
Now that we’ve identified outlier loci, it is useful to be able to
identify how much they correlate to each environmental variable. To do
so, we can run a correlation test of each SNP using
rda_cor() and summarize results in a customizable table
using rda_table().
# Extract genotypes for outlier SNPs
rda_snps <- rda_sig_p$rda_snps
rda_gen <- gen[, rda_snps]
# Run correlation test
cor_df <- rda_cor(rda_gen, env)
# Make a table from these results (displaying only the first 5 rows):
rda_table(cor_df, nrow = 5)| r | p | snp | var |
|---|---|---|---|
| 0.27 | 0.02 | Locus_125 | CA_rPCA3 |
| 0.39 | 0.00 | Locus_166 | CA_rPCA3 |
| 0.29 | 0.01 | Locus_249 | CA_rPCA2 |
| -0.24 | 0.03 | Locus_263 | CA_rPCA1 |
| 0.27 | 0.02 | Locus_263 | CA_rPCA3 |
# Order by the strength of the correlation
rda_table(cor_df, order = TRUE, nrow = 5)| r | p | snp | var |
|---|---|---|---|
| -0.49 | 0 | Locus_2947 | CA_rPCA2 |
| -0.43 | 0 | Locus_1278 | CA_rPCA2 |
| -0.42 | 0 | Locus_2338 | CA_rPCA3 |
| -0.42 | 0 | Locus_786 | CA_rPCA2 |
| -0.42 | 0 | Locus_1318 | CA_rPCA2 |
# Only retain the top variable for each SNP based on the strength of the correlation
rda_table(cor_df, top = TRUE, nrow = 5)| r | p | snp | var |
|---|---|---|---|
| 0.27 | 0.02 | Locus_125 | CA_rPCA3 |
| 0.39 | 0.00 | Locus_166 | CA_rPCA3 |
| 0.29 | 0.01 | Locus_249 | CA_rPCA2 |
| 0.27 | 0.02 | Locus_263 | CA_rPCA3 |
| 0.28 | 0.01 | Locus_312 | CA_rPCA2 |
# Display results for only one environmental variable
rda_table(cor_df, var = "CA_rPCA2", nrow = 5)| r | p | snp | var |
|---|---|---|---|
| 0.29 | 0.01 | Locus_249 | CA_rPCA2 |
| 0.28 | 0.01 | Locus_312 | CA_rPCA2 |
| 0.27 | 0.02 | Locus_404 | CA_rPCA2 |
| 0.34 | 0.00 | Locus_517 | CA_rPCA2 |
| 0.23 | 0.04 | Locus_612 | CA_rPCA2 |
Running RDA with rda_do_everything()
The algatr package also has an option to run all of the above
functionality in a single function, rda_do_everything().
This function is similar in structure to rda_run(), but
also performs imputation (impute option), can generate
figures, the correlation test table (cortest option;
default is TRUE), and variance partitioning (varpart
option; default is FALSE). Please be aware that the
do_everything() functions are meant to be exploratory. We
do not recommend their use for final analyses unless certain they are
properly parameterized.
The resulting object from rda_do_everything()
contains:
Candidate SNPs:
rda_snpsData frame of environmental associations with outlier SNPs:
cor_dfRDA model specifics:
rda_modOutlier test results:
rda_outlier_testRelevant R-squared values:
rsqANOVA results:
anovaList of all p-values:
pvaluesVariance partitioning results:
varpart
Let’s first run a simple RDA, imputing to the median, with variable selection and the p-value outlier method:
results <- rda_do_everything(liz_vcf, CA_env, liz_coords,
impute = "simple",
correctGEO = FALSE,
correctPC = FALSE,
outlier_method = "p",
sig = 0.05,
p_adj = "fdr",
cortest = TRUE,
varpart = FALSE
)
| r | p | snp | var |
|---|---|---|---|
| -0.49 | 0 | Locus_2947 | CA_rPCA2 |
| -0.43 | 0 | Locus_1278 | CA_rPCA2 |
| -0.42 | 0 | Locus_2338 | CA_rPCA3 |
| -0.42 | 0 | Locus_786 | CA_rPCA2 |
| -0.42 | 0 | Locus_1318 | CA_rPCA2 |
| -0.42 | 0 | Locus_771 | CA_rPCA2 |
| -0.41 | 0 | Locus_1374 | CA_rPCA2 |
| 0.40 | 0 | Locus_636 | CA_rPCA3 |
| 0.39 | 0 | Locus_1263 | CA_rPCA2 |
| 0.39 | 0 | Locus_166 | CA_rPCA3 |
Now, let’s run a partial RDA, imputing based on 3 sNMF clusters,
correcting for both geography and structure, and instead using the
Z-score method to detect outlier loci, including variance partitioning.
We’ll keep the number of PCs for population structure set to the
default, which is three axes (nPC):
results <- rda_do_everything(liz_vcf, CA_env, liz_coords,
impute = "structure",
K_impute = 3,
model = "full",
correctGEO = TRUE,
correctPC = TRUE,
nPC = 3,
varpart = TRUE,
outlier_method = "z",
z = 3
)
| Variance partitioning | ||||||
| Model | Inertia | R2 | Adjusted R2 | p (>F) | Prop. of explainable variance | Prop. of total variance |
|---|---|---|---|---|---|---|
| Full model | 70.50 | 0.44 | 0.35 | 0.00 | 1.00 | 0.44 |
| Pure enviro. model | 4.80 | 0.03 | 0.01 | 0.02 | 0.07 | 0.03 |
| Pure pop. structure model | 29.48 | 0.18 | 0.16 | 0.00 | 0.42 | 0.18 |
| Pure geography model | 5.43 | 0.03 | 0.01 | 0.00 | 0.08 | 0.03 |
| Confounded variance | 30.80 |
|
|
|
0.44 | 0.19 |
| Total unexplained variance | 89.95 |
|
|
|
|
0.56 |
| Total inertia | 160.45 |
|
|
|
|
1.00 |
| r | p | snp | var |
|---|---|---|---|
| -0.42 | 0.00 | Locus_2338 | CA_rPCA3 |
| -0.35 | 0.00 | Locus_2767 | CA_rPCA1 |
| 0.31 | 0.01 | Locus_2665 | CA_rPCA2 |
| 0.29 | 0.01 | Locus_2053 | CA_rPCA1 |
| -0.29 | 0.01 | Locus_1005 | CA_rPCA1 |
| -0.29 | 0.01 | Locus_1005 | CA_rPCA1 |
| 0.29 | 0.01 | Locus_3058 | CA_rPCA3 |
| 0.28 | 0.01 | Locus_1660 | CA_rPCA2 |
| 0.27 | 0.01 | Locus_469 | CA_rPCA3 |
| 0.27 | 0.02 | Locus_2995 | CA_rPCA2 |
Additional documentation and citations
| Citation/URL | Details | |
|---|---|---|
| Associated literature | Capblancq & Forester 2021; Github repository for code | Paper describing RDA methodology; also provides walkthrough and
description of rdadapt() function |
| Associated literature | Capblancq et al. 2018 | Description of p-value method of detecting outlier loci |
| R package documentation | Oksanen et al. 2022. | algatr uses the rda() and ordiR2step()
functions in the vegan package |
| Associated literature | Forester et al. 2018 | Review of genotype-environment association methods |
| Associated tutorial | Forester et al. | Walkthrough of performing an RDA with the Z-scores outlier detection method |
| Associated vignette | vignette("intro-vegan") |
Introduction to ordination using the vegan package (including using
the rda() function |
| Associated vignette | vignette("partitioning") |
Introduction to variance partitioning in vegan |